B. Papp szerk.: Studia Botanica Hungarica 39. 2008 (Budapest, 2008)

Erzberger, P.; Schröder, W.: The genus Schistidium (Grimmiaceae, Musci) in Hungary

Costa and leaf margins smooth. Lamina smooth, in lower part unistratose, in upper part with bistratose spots. Upper lamina cells rounded, 6-9 urn wide, lower cells wider and slightly sinuose, basal marginal cells sometimes subhyaline, with slightly thickened cross-walls. K+ red. Sporophytes frequent, mostly shallowly immersed. Urn orange to reddish-brown, often finely striolate, widest below middle, length/width ratio (1.3-) 1.4-1.6-1.8 (BLOM 1996), in Hungarian material (0.94-) 1.34 ±0.04(-1.93), n = 91. Exothecial cells elongate, especially in basal part, at least some up to 50 urn (often up to 80 urn) long, arranged in a regular pattern. Stomata absent. Peristome teeth 220-340 urn, orange, in central and upper parts mostly strongly perforated to cribrose. Variability: The exothecial cells in the lower part of the urn sometimes only reach (50-)55 urn, but in other samples they attain 75-80 urn. The brown, dense cushions with hair-points only around the sporophytes give this plant a very characteristic appearance. However, some forms of S. crassipilum can look quite similar, and in that case the strongly elongate exothecial cells (Fig. 5G, H, compare with Fig. 131, J), and the shorter and wider urn of S. brunnescens are diagnostic. Confusion could also occur with forms of S. belveticum lacking the typical black colour, but again the pattern of exothecial cells is quite different. For the difference between subsp. brunnescens and subsp. griseum see the note under the latter. Habitat: S. brunnescens subsp. brunnescens grows nearly exclusively on exposed calcareous rocks (limestone, dolomite), and is a very typical member of the bryophyte vegetation in calcareous grasslands. It was found associated with the lichen Fulgensiafulgens (det. V. Otte). Associated bryophytes: Bryum argenteum, Didymodon rigidulus, D. vinealis, Grimmia dissimulata, G. orbicularis, G. pulvinata, Grimmia tergestina, Pseudocrossidium revolutum, Pseudoleskeella catenulata, Scbistidium crassipilum, S. belveticum, Tortella inclinata, Tortula crinita, Tricbostomum sp. Vertical distribution: 150-780 m a.s.l. Selected specimens examined: Aggtelek Karst: 7589/1 Comit. Gömör. In rupibus calc. montis "Baradla-tetö" prope pag. Aggtelek, 350 m, 03.06.1928, leg. A. Boros, BP 111757; Bükk Mts: 7988/3 Comit. Borsod. In rupibus calcar. montis Bélkö prope Bélapátfalva, 500-780 m, 03.09.1959, leg. Á. Boros, BP 110944; 8088/2 Lehmschieferfelsen an der Straßenböschung zwischen Bükkzsérc und Felsőtárkány, ca 400 m, mit S. crassipilum, 27.03. 2008, leg. P. Erzberger and T. Poes, herb. Erzberger, (B) 12886; Tarna Region: 7986/1 Glaukonit-Sandsteinfelsen "Noé szöllöje" im Ort Istenmezeje, 230 m, 26.03.2008, leg. P. Erzberger and T. Poes, herb. Erzberger, (B) 12861 ; Cserhát Hills: 8181/4 Comit. Pest. In rupibus dolomit, apricis montis Vár-hegy prope Csővár, 350 m, 06.05.1953, leg. A. Boros, BP 111748; Gödöllő Hills: 8381/3 Comit. Pest. In monte Somlyó prope Fóth, 21.05.1901, leg. Á. Degen, BP 37650; Pilis Mts: 8380/3 Comit. Pest. In rupibus dolomiticis merid. montis Nagykevély prope pag. Pilisborosjenő, 500 m, 25.03.1947, leg. Á. Boros, BP 111750; Hung, centr., com. Pest. In rupibus calcareis supra pag. Pomáz, 31.03.1946, leg. L. Vajda, EGR; Buda Mts: 8380/3 Budakalász, comit. Pest, in monte "Monalovác" ad saxa calcarea, 250 m, III. 1911, leg. J. Szurák, revid. M. Péterfi, BP 5645; Gerecse Mts: 8477/2 Gyermely,