Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 92. (Budapest 2000)

Roháček, J. ; Papp, L.: Crumomyia microps sp. n. from Austria and notes on other cavernicolous Crumomyia species (Diptera: Sphaeroceridae)

Discussion - The new species is a member of the Crumomyia notabilis-gvoup sensu NORRBOM & KIM (1985). Judging from the highly similar male genitalia it is the closest relative of Crumomyia notabilis (COLLIN, 1902) and, consequently, its sister-species. Other closely allied species undoubtedly are: C. absoloni (BEZZI, 1914), C. rohaceki NORRBOM et KlM, 1985 and C. zuskai (ROHÁCEK, 1976). All these species share similar microtomentose pattern on gena, similar construction of distiphallus with a pair of small dorsal hooks and postgonite with simple lateral outline but with internal tooth or lobe. Interestingly, all these species can live also (if not only) in caves. In addition, C. hungarica (DUDA, 1938) should also be taken into consideration although its membership in the above assemblage is not certain because the species is only known from the female holotype (see PAPP 1979). Owing to strongly reduced eyes, C. microps sp. n. externally most resembles the cavernicolous species C. absoloni and C. hungarica. However, both these species differ from C. microps sp. n. in having strongly shortened wings (cf. PAPP 1979, Figs 5, 6), not to mention different gonostylus (apically pointed), postgonite (apically somewhat notched) and female 8th tergum (medially undivided) of C. absoloni (cf. NORRBOM & KlM 1985, Figs 121,122,124) and only one orbital seta, enlarged prescutellar dorsocentral seta and pale legs of C. hungarica. As noted above, the closest ally of C. microps sp. n., viz. C. notabilis, has very similar male genitalia but differs in a number of external features, e.g. in having darker (almost black) body, shorter and thicker macrosetae on head and thorax, shorter cilia on arista, usually additional setulae on scutellum, shorter and thicker legs, darker clouded cross-veins, heavily sclerotized preabdominal terga and, most distinctly, large eyes (cf. NORRBOM & KlM 1985, Fig. 127) and mesopleural microtomentose pattern restricted to dorsal marginal area (see ROHÁCEK 1976, Fig. 2). However, the validity of C. microps sp. n. can also be supported by several characters of the male and female terminalia, particularly by the epandrium without enlarged setae, apically rounded gonostylus (Fig. 7), strongly bent epiphallus, smaller paired hooks on distiphallus (all on Fig. 12) and larger inner tooth of postgonite (Fig. 8) and by the medially divided 8th female tergum. Biology - All but one type specimens of C. microps sp. n. were found in an Alpine stalactite cave during a-year period (precise dates of collection unknown) at unusually high altitude (2,320 m). The flies occurred here in community with Crumomyia cavernicola oculea ssp. n. (described below), which was probably caught in the entrance zone of the cave, and the very rare - cavernicolous but not troglobiont - Eccoptomera sanmartini CZERNY, 1924 (described from San Martino di Castrozza and known only from gr. du Lichen, Onnion, Haute-Savoie, see PAPP 1982). One paratype was found in a nearby ice cave at an altitude of 2,180 m. Distribution - Hitherto, the species is only known from the Obstans caves in the Karnische Alpen Mts (Austria).