Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 92. (Budapest 2000)

Roháček, J. ; Papp, L.: Crumomyia microps sp. n. from Austria and notes on other cavernicolous Crumomyia species (Diptera: Sphaeroceridae)

TAXONOMY OF THE CRUMOMYIA PARENTELA COMPLEX Two females and three males of a Crumomyia species found in the Obstans caves are externally highly similar to the montane Crumomyia parentela alpicola (ROHÁCEK in TROGER et ROHÁCEK 1980) but proved to have somewhat different genitalia, resembling those of C. parentela cavernicola (PAPP et ROHÁCEK, 1983). A female Crumomyia parentela alpicola found also in the Obstans cave (Austria: Osttirol, Kamische Alpen, Obstans Tropfsteinhöhle, 2320 m, 20.9.1998-15.9.1999, Kahlen leg., in coll. A. KOFLER, Lienz) confirms our presupposition that both C. parentela and C. cavernicola occur sympatrically in the Karnischen Alpen and have therefore be treated as separate species. The female examined has postabdomen with characteristically long, anteriorly incised but medially undivided 8th sternum. It was captured in the same cave as three males of C. cavernicola oculea ssp. n. (described below). This fact initiated a more thorough comparative study of all taxa belonging to the C. parentela complex, which resulted in finding that the differences in genitalic structures are invariable and enable us to split this complex of subspecies into two groups which are classified here as separate species. Each of these species include two subspecies: one troglobiont with strongly reduced eyes, narrowed thorax and prolonged legs, and the other free-living (or at most troglophilous) one with eyes, thorax and legs normal (for a classification of cave dwelling flies see below). The taxonomical results can be summarized as follows: Crumomyia parentela (SÉGUY, 1963) ssp. parentela (SÉGUY, 1963) - a troglophilous subspecies known from two caves in Serbia (see NORRBOM & KlM 1985). ssp. alpicola (ROHÁCEK in TROGER et ROHÁCEK, 1980) - a montane subspecies widespread in Alps and Carpathians, also known from caves in Slovakia (see ROHÁCEK 1991). Crumomyia cavernicola (PAPP et ROHÁCEK, 1983), stat. n. ssp. cavernicola (PAPP et ROHÁCEK, 1983) - a troglobiont subspecies described from St. Vincent cave in France. ssp. oculea ssp. n. - a troglophilous subspecies from Obstans caves in Austria. It supposedly also occurs outside the caves and can thus be expected elsewhere in Alps. For diagnosis and type material see below.