S. Mahunka szerk.: Folia Entomologica Hungarica 63. (Budapest, 2002)

lobe, vein CuAl terminus lobate, vein CuA2 terminus tailed. Dorsal surface: fore wing ground colour marine blue with very narrow black outer margin and brown distal fringes, hind wing similar to fore wing but with black tail at vein CuA2 terminus plus tomal and anal margin lobes and gleaming blue antemarginal stripes. Ventral surface: fore wing ground colour warm brown with median and submarginal vertical row of gleaming blue intercellular lines, antemarginal area with faint lighter blue vertical stripe; hind wing colouration as on fore wing but with submedian pattern of intercellular lines from cell Sc+Rl to cell CuAl, a postmedian line formed by lineal intercellular pattern, a submarginal row of shiny antemarginal intercellular lines from cell Ml to tornus, a vivid blue marginal line, and tornal and inner marginal lobes black. Body gleaming blue dorsally, brown ventrally. Genitalia: Not available for study (abdomen lacking). Male: Not known. Etymology - A freely formed noun, gender masculine, from the name of lepidopterist Bernard d'Abrera, who was the first to recognize the species and point out its relationship. Note - d'Abrera (1995,1. c.) noted that the specimen herein selected as holotype was related to M. danaus and figured its dorsal and ventral surfaces. He also indicated that the specimen is a male. Knowing the congeners well, I consider the specimen to be a female because the distal margins of the fore wings of male Margaritheclus are straight and possess an androconial cluster, and their fore wing ventral surface do not show pattern or have different ground colours from those of the hind wing. DISCUSSION Placement in Pseudolycaena Wallengren - The original placement of the genus's type species is erroneous as the type species of Pseudolycaena Wallengren, 1858 is Papilio marsyas Linnaeus, 1758 by original designation. P. marsyas possesses qualitatively different wingshape, dorsal wing colouration, and ventral pattern (cf. d'Abrera 1995: 1112-1113), as well as distinct male genitalia (cf. Clench 1964: fig. 2) and female genitalia (a long tube with a flat posterior plate showing two pointed symmetrical lateral processes). Moreover Pseudolycaena males possess a ventral androconial pouch in the middle of the cell CuA2 of the hind wings, what I consider as a homolgous trait of the ventral androconial pouch of Denivia and certain species of Theritas Hübner, 1819 (Bálint in prep.). It is thus not possible to lump P. marsyas and M. danaus (and its congeners described herein) in the same cluster of taxa as they are not even close sister lineages within the Eumaeini; therefore the original placement of the Felders did not reflect monophyly.