S. Mahunka szerk.: Folia Entomologica Hungarica 63. (Budapest, 2002)

Placement in Tolmides Group of Thecla sensu Draudt - This group was formed for the nominal taxa Thecla tolmides (Felder et Felder, 1865), Th. tityrus (Felder et Felder, 1865), Th. appula (Hewitson, 1867) and Th. danaus. This cluster of taxa is also not monophyletic. All its members possess quantitatively distinct wingshape, qualitatively different male dorsal androconia (if present), ventral wing pattern, and genital structures. The taxon tolmides was transferred to genus Egides Johnson, Kruse et Kroenlein, 1997 (type species: Pseudolycaena aegides Felder et Felder, 1865, by original designation), which is erroneous as the type species of the genus basically differs from tolmides in wingshape, colouration and pattern. The taxon tolmides is closely related to Laothus, but its ventral wing pattern and female genitalia are supralimital. It has been already placed in a distinct group within Eumaeini (Luis-Martinez et al. 1995: 127). Additionally, Egides can be considered monophyletic with Micandra Schatz, 1888 (type species: Pseudolycaena platyptera Felder et Felder, 1865, by monotypy) and tolmides was regarded as a species of Micandra by Clench (1971: 2) and Robbins (1986: 150). Pseudolycaena tityrus Felder et Felder, 1865 cannot be monophyletic with M. danaus; its males possess a dorsal androconial cluster filling the discal cell completely, the wingshape is different, as is the ventral wing pattern. The male possesses genital brush organ, and lacks a lateral projection on the valva, while in the female the genital ductus is a very long simple tube, reminiscent of Pseudolycanea. The species tityrus was selected as type species of Balintus d'Abrera, 2001 (p. 195), a genus he considered as monotypic. I have found a sister species of B. tityrus from Colombia in recently collected material. Regarding Thecla appula, this taxon was transferred to Parrhasius Hübner, 1819 (type species: Papilio polibetes Stoll, 1781) by Nicolay (1979: 27) and belongs to the cluster of genera including Iaspis Kaye, 1904, Michaelus Nicolay, 1979 and Panthiades Hübner, 1819, therefore M. danaus cannot be monophyletic with P. appula. Placement in Havila Group of Thecla sensu Draudt - This group was formed for the nominal taxa Thecla adamsi (Druce, 1907), Th. havila (Hewitson, 1867), Th. celelata (Hewitson, 1867) and Th. margaritacea (Draudt, 1919). It has been already mentioned that the last taxon is the male of M. danaus according to Goodson (1946: 270). I here transfer the taxon adamsi to Denivia Johnson, 1992 (type species Thecla deniva Hewitson, 1877) thus, Denivia adamsi (Druce, 1907), new combination, since the holotype male possesses the ventral androconial pouch along hind wing vein 1A+2A. I consider this trait as an apomorphic for Denivia (Bálint in prep.). Th. havila is a supralimital member of the genus Atlides Hübner, 1819 (type species: Papilio halesus Cramer, 1777) as its male has the wingshape, androconial cluster, and ventral abdominal colouration typical of Atlides; therefore I transfer the species to that genus: Atlides havila (Hewitson, 1865), new combi-