O. Gy. Dely szerk.: Vertebrata Hungarica 23. (Budapest, 1989)

Topál, Gy.: Tertiary and Early Quaternary remains of Corynorhinus and Plecotus from Hungary (Mammalia, Chiroptera) 33-55. o.

Among the available tooth-row lengths, the C-M3/ alveolar length is slightly greater than in P. townsendii

, while the C-P4/ alveolar length is smaller than that of the recent animal. Other measurable tooth-row lengths overlap extensively in the two species (see Table 1), As regards the individual upper teeth, P2/ basal length is clearly greater than in P. town sendii, though the width of this tooth is equal in both species. On the contrary, the P4/ basal length of P. crassidens is appreciably smaller than even that of the smallest P. townsendii studied. Otherwise, all measurements but the length of M2/ seems to exceed those of P. townsendii. In mandibular morphology the most important difference between P. crassidens and P. townsendii lies in the shape of the basal portion of the angular process on the buccal side. It has a convexity and a sharp ridge in P. crassidens (anatomically corresponds to the linea masseterica in the dog), while in P. townsendii the angular process is more intensively concavely bent outwards in occlusal view, and without a ridge (see Plate I, Figs 3, 5). The height of the mandibular body under M/l exceeds that of P. townsendii

, otherwise the mandible does not seem to differ in the two species. As regards the available lower tooth-row length measurements, these overlap to various extent in the two species. Basal length and especially basal width, thus the crown area of P/2 is clearly greater than in P. townsendii, contrarily, the basal width of P/3 is much smaller in P. crassidens (while its basal length is equal) so this premolar is definitely narrowed, that is, specialized in the species P. crassidens (see Plate II, Figs 3, 5). See also RABEDER (op. cit. Table VI, Fig. 53). The basal length and width of P/4 do not differ in the two species and there are no differences in other tooth measurements, except in M/l talonid width which is smaller in P. crassidens, and in M/3 talonid width which is wider in P. crassidens than in the recent species. The tooth measurements of the holotype of P. crassidens (as shown above) are greater than those of the extant species. It has also longer M/3 with wider trigonid and talonid. The species P. crassidens is a more evolved, over specialized form of a more or less separate lineage within the subgenus Corynorhinus. Plecotus (Corynorhinus) atavus Topái, 1988 At present, this is one of the earliest known plecotine in the Holarctic. It comes from the Upper Miocene (MN 13 zone), Polgárdi Locality 4. A rather primitive plecotine (Corynorhinusl bat with clearly two-rooted P/4 and partly ancestral, partly intermediate characters as compared to Plecotus (Corynorhinus) crassidens Kormos, 1930 and Plecotus (Coryn

o

rhinus) townsendii pallescens Miller, 1897. Holotype : Right maxillary fragment (Plate I, Figs 7, 8) with P4/-M1/-M2/ and alveoli of I1/-I2/, C, M3/. Register No. V. 15228 (VT. 138). Deposited in the Palaeovertebrate Collection, Hungarian Geological Institute, Budapest. Measurements of the holotype: DAPE 1. 90, MHP 1. 50, MHM 1. 22, AB 0. 55, UCMA 5. 35, UPM 4. 23, UCPA 2.31, UMMA 3. 44, UCLA 0. 72, UCWA 0. 72, UPFL 1. 29, UPFW 1. 33, UMOL 1. 45, UMOW 1. 72, UMTL 1. 36, UMTW 1.74. Other material and a selected paratype: V. 15230 right maxillary fragment with P2/-damaged P4/-Ml/-M2/-damaged M3/; Reg. No. V. 15231 right maxillary fragment with P4/, M2/-M3/ and alveolus of Ml/; Reg. No. V. 15232 right detached Ml/; Reg. No. V. 15234 right maxillary fragment with P4/, M2/ and alveoli of II/P2/, Ml/, M3/; Reg. No. V. 15235 slightly fragmentary (with missing angular process) right mandible with alveoli of I/l-M/3; Reg. No. V. 15233 right mandibular fragment with C-P/2-P/3-P/4-M/l-M/2 and alveoli of I/1-I/3; selected paratype Reg. No. 5. 15229 (Plate II, Figs 1, 2) left mandible with P/4 and alveoli of I/l-P/3, M/l-M/3. All in the Palaeovertebrate Collection, Hungarian Geological Institute, Budapest. Measurements of the selected paratype: ML 11. 50, LCMA 6.07, LCPA 2.25, LLMA 3.79, H MUM 1.68, HNBM 1.50, HCP 3.56. Measurements: (including-those of holotype and selected paratype) minimum and maximum (sample size and mean in parentheses). DAPE 1.81-1.90 (2; 1.855); MHP 1.361. 50 (3; 1. 436); MHM 1. 00-1. 36 (4; 1. 177); AB 0. 5-0. 68 (3; 0. 626); UCMA 5.08-5. 35 (2; 5.215); UPM 4. 19-4.23 (2; 4.210); UPMA 3.95-4. 23 (4; 4.092); UCPA 2.03-2. 31 (2; 2.170); •