O. Gy. Dely szerk.: Vertebrata Hungarica 23. (Budapest, 1989)

Topál, Gy.: Tertiary and Early Quaternary remains of Corynorhinus and Plecotus from Hungary (Mammalia, Chiroptera) 33-55. o.

UMM 3. 46; UMMA 3. 08-3. 44 (4; 3. 240); UCLA 0. 68-0. 72 (2; 0. 70); UCWA 0. 63-0. 72 (2; 0. 675); UPTL 0. 50; UPTW 0. 55; UPFL 1. 18-1. 29 (4; 1. 245); UPFW 1. 18- 1. 36 (4; 1. 27 5); UMOL 1.40-1.45 (3; 1.433); U MOW 1. 65- 1.77 (3; 1.713); UMTL 1.29- 1.45 (4; 1.365); UMTW 1. 65-1. 79 {4; 1. 7 37); UTML 0. 81-0. 81 (2; 0.81); UTMW 1. 68- 1. 72 (2; 1. 70); ML 10.65-11.50 (2; 11.075); LCMA 5.85-6.07 (2; 5.955); LPM 2. 18; LCPA 1. 95-2. 25 (3; 2.10); LMMA 3.66- 3.79 (2; 3.725); LCL 0. 50; LCW0.86; LPTL 0. 45; LPTW0.50; LPFL 0. 680.77 (2; 0.725); LPFW 0. 65-0. 72 (2; 0.685); LMOL 1.36; LMOW 0.95; LMTL 1. 365; LMTW 0.95; HMUM 1.45-1. 68 (3; 1. 556); H MB M 1.31-1.50 (2; 1.405); HCP 3.45-3. 56 (2; 3. 450). Description and comparisons: No doubts there may be differences in the morphology of the rostrum of P. atavus and the already known P. crassidens and P. town sendii used for comparison in this study. It must be admitted that a number of characters are not properly seen in P. atavus because of its rather fragmentary state. The lacrimal area is smooth as in the other species studied. Regarding the dor so-median rostral depression the species is most probably closer to P. crassidens than to P. townsendii. P. atavus has a medium-sized anteorbital foramen as compared to the other species. It is placed mostly above the P4/ and partly above the Ml/ area. The distance from the posterior margin of anteorbital foramen to the foremost tip of the premaxilla is intermediate between that of P. crassidens and P. townsendii. The anterior palatal emargination was probably greater, first of all deeper than even that of P. crassidens. It extends back to the connecting line between P2/-P2/ alveolar centers, while in Beremend P. crassidens it is somewhat less deep. The maxillary height above the anterior root of P4/ is much lower in P. atavus than in P. crassidens and in P. townsendii (see Table 1). P. atavus and P. townsendii are virtually identical in the maxillary height above M2/ or even some Miocene specimens have greater values than P. townsendii has, so these are greater than in P. crassidens. Regarding the mean values of the upper tooth-row lengths (C-M3/ alveolar length and M1/-M3/ length) the Miocene species is identical with P. townsendii, thus both are smaller than P. crassidens. On the contrary, however, in C-P4/ alveolar length and P4/M3/ length P. atavus and P. townsendii are equally greater than P. crassidens. Though the upper incisors are missing, one can infer from the alveoli that P. crassidens had definitely greater II/ as compared with 12/ than P. atavus

. The difference between the two incisors is much less pronounced in P. townsendii. The length of the alveole of the upper canine in P. crassidens proved to be greater with no overlap with the new species. One can infer a laterally relatively more pressed, antero-posteriorly more elongated root of canine than that of P. atavus. Unfortunately, until now, the upper canine of P. crassidens, nor that of P. atavus has been found. The bulk of the P2/ of P. atavus is comparatively great and" steadily decreasing in basal length from the Miocene P. atavus through the Pleistocene P. crassidens to the recent species. P2/ basal width, however, seems to differ in the two fossil species (one specimen for each). In P. townsendii the range includes all fossil values (see Table 1). The available material of P4/ basal lengths does not overlap between P. atavus and P. townsendii, and at the same time both have greater values of this character than P. crassidens does. Besides, all three species show overlap in P4/ widths. These indicate a pronounced antero-posterior shortening of P4/ in P. crassidens, while P. atavus is less specialized in this respect. There is a characteristic difference in the morphology of the crown of P4/ between the species studied. The anterior edge of the crown has an outer anterior lobe and a deep incision for keeping the posterior portion of the small premolar in P. atavus. These are missing in P. crassidens where the anterior border of P4/ is very slightly pressed by P2/, so it is almost straight without an undulation shown by the former species (see also RABEDER op. cit. Plate II, Fig. 8a). In P. townsendii the anterior border is straight or with a slight lagune. The lingual portion of Ml/ and M2/ is bulky in P. crassidens but slender in P. atavus. M3/ is less specialized (less shortened) in P_

I

_crassidejis_, intermediate in P. atavus and variable but generally most specialized (shortened) in the available specimens of P. townsendi

i. The talon is more or less distinct from the trigon by a narrowing neck in the fossil species, there is no such narrowness in P. crassidens. The length of the mandible shows overlap in the compared three species, the smallest specimens being found in P. townsendii and the greatest in P. atavu

s. The foramen mentale is situated under the portion between C and P/2, not under the two small premolars