L. Hably szerk.: Studia Botanica Hungarica 20. 1988 (Budapest, 1988)

Hably, Lilla: Egerian fossil flora from Kesztölc, NW Hungary

1980 Platanus neptuni (Ett.) Bűzek, Holy et Kvacek; Habiy, p. 300, Pl. 1, Figs 1-8; Pl. 2, Figs 1-9; Pl. 3, Figs 1-6; Pl. 4, Figs 1-5; Pl. 5, Figs 1-6; Pl. 6, Figs 1-4; Pl. 7, Figs 1-4; Pl. 8, Figs 1-4; Pl. 9, Figs 1-4; Pl. 10, Figs 1-4. 1982 Platanus neptuni (Ett.) Buzek, Holy.et Kvacek; Hably, Pl. 1, Figs 4, 6, 9, 10, 11; Pl. 2, Figs 1-2; Pl. 5, Figs 1-6; Pl. 6, Figs 1-6; Pl. 7, Figs 2-4; Pl. 8, Fig. 1. 1985 Platanus neptuni (Ett.) BÛzek, Holy­et Kvacek; Hably, p. 101, Pl. 17, Fig. 6; Pl. 18, Figs 1-6; Pl. 19, Figs 1-5; Pl. 20, Figs 1-4, Pl. 21, Figs 1-3. Material: BP: 85. 274. 1., 85. 317. 1., 85. 325. 1., 85. 326. 1. MÁFI: BK­1107, BK-1109, BK-1117, BK-1120, BK-1121, BK-1122, BK­1127, BK-1141, BK-1142, BK-2465, BK-2915 The length of the leaves 3. 0-9. 5 cm, their width between 0. 8-2. 8 cm. Some of them are tanged, while others are simple leaves without tangs. Their shape is generally elongated oval, in some cases elongated obovate. The apex and the base are acute, the basis is typical­ly asymmetrical in almost all specimens. The midvein is strong, the venation is camptodro­mous. Quite often, intermediate veins among the secondary veins can be observed, branch­ing out from the midvein. The margin of the leaf is entire in the lower third, occasionally in the lower half, whereas in the apical phase it is toothed. The teeth are fairly characteristic, their apex is blunt, their apical side is essentially shorter than that of the basal one. The apical side is straight or concave, the basal side is slightly concave. The sinuses between the teeth are rounded. The species is widely distributed in the Egerian stage of Hungary (HABLY 1980, 1982, 1983, 1985b and in press), and is equally present in the Lower Oligocène (HABLY 1979) and the Lower Miocene (HABLY 1985a). In Europe, it is known from the Eocene till the Karpatian from a number of localities. It is a palaeotropical element, a member of the subtropical rain­forests, present mainly in the period of climate improvements. Probably it used to consti­tute extensive forests in the Egerian Stage of Hungary, which is indicated by its general and mass appearance on the Egerian localities. Platanus fraxinifolia (John, et Gilm. ) Walther Pl. VIII, Figs 1-2, 4, Pl. VII, Fig. 3. 1921 Dewalquea hibernica Johnson et Gilmore; Johnson et Gilmore, p. 325, PI. 11, Figs 1-3; PI. 12, Figs 1, 2, 5, 11; Text-fig. 1. 1921 Dewalquea fraxinifolia Johnson et Gilmore; Johnson et Gilmore, p. 327, PI. 11, Figs 4-11; PI. 12, Fig. 3; Text-fig. 2. 1921 Dewalquea denticulata Johnson et Gilmore; Johnson et Gilmore, p. 327, PI. 11, Figs 12-14; PI. 12, Fig. 4; Text-fig. 3. 1963 Dicotylophyllum bellum (Heer) Sew. et Conw. ; Koch, p. 66, PI. 35, PI. 36, PI. 37; Text-fig. 24. ? 1921 Juglandiphyllum denticulatum (Heer) Koch; Koch, p. 42, PI. 20, Fig. 3; PI. 11, PI. 12, PI. 13, PI. 14. 1985 Platanus fraxinifolia (John, et Gilm.) Walther; Mai et Walther, p. 55, PI. 13, Figs 1-8; PI. 9/3-6, p. 195, PI. 13/1-2, p. 203. Material: BP: 85. 361. 1., 85. 369. 1. MÁFI: BK­1104, BK-1120, BK-1127, BK-2462, BK-2464, BK-2492 The leaf is 6. 5-10; 5 cm long, found with leaflets of 1. 7-2. 5 cm width. The leaflets are strongly elongated ovate or ellipsoid, the apex and the basis is acute. The venation is campto­dromous. Intermediate veins branch out from the midvein between the secondary veins. The margin of the leaf is toothed. The teeth are rather similar to those of P. neptuni . In the ba­sal part, the margin is entire, similar to the above species. There was no complete leaf occurring at the Kesztölc site, still on the basis of some further finds of Egerian flora so far unpublished, the remains can be assigned to this species. The shape of the leaves and the character of the basis is essentially different from those of P. neptuni , though in respect of the character of the denticulation, they are very close to each other. In some dubious cases it is really hard to separate them. It can be supposed that all these remains are to be considered as one species of very wide range of variation. This hypothesis might be supported by the fact that both forms coexist in most of the Egerian lo-

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