L. Hably szerk.: Studia Botanica Hungarica 20. 1988 (Budapest, 1988)

Hably, Lilla: Egerian fossil flora from Kesztölc, NW Hungary

calities in Hungary. At the same time, no P. fraxinifolia was found at the Kiscellian local­ities or at Ipolytarnóc, though these localities were fairly rich in P. neptuni . Fagaceae Castanopsis (D. Don. ) Spach cf. Castanopsis toscana (Bandulska) Kräusel et Weyland Pl. VIII, Figs 5-6; Pl. IX, Fig. 1. 1931 Tristania toscana Band. ; Bandulska, p. 668, Pl. 40, Figs 9-11. 1954 Castanopsis toscana (Band.) Kr. et Weyl. ; Kräusel et Weyland, p. 135, Text-fig. 14, Pl. 29, Figs 3-9. 1971 L. sensu Ferguson; Ferguson, pp. 207-216, Text-fig. 37, Pl. 34-36. 1976 Castanopsis toscana (Band.) Kr. et Weyl. ; Knobloch et Kvaiiek, p. 39, Pl. 4, Fig. 1; Pl. 16, Figs 10-11; Pl. 17, Fig. 16; Pl. 20, Figs 11, 15. Material: BP: 85. 275. 2., 85. 282. 1., 85. 335. 1., 85. 337. 1., 85. 338. 1., 85. 340. 1., 85. 341. 1., 85. 342. 1., 85. 344. 1., 85. 347. 1., 85. 348. 1., 85. 350. 1., 85.'354. 1., 85. 356. 1. MÁFI: BK­1126, BK-2455, BK-2470, BK-2474, BK-2497, BK-2932, BK-2933, BK-2934, BK-2935, BK-2936, BK-2937, BK-2938, BK-2939, BK-2940 The length of the leaves is 3. 7-12. 3 cm, their width varies between 1. 4 and 2. 8 cm. Their shape is elongated ellipsoidal, elongated or ovate. The apex of the leaf is acute or at­tenuate, the base is acute or rounded. The margin of the leaf is entire, the venation is cam­ptodromous. Some of the leaves are petiolated, while others are without petiole. The species is represented in the floral assemblage in very high number. KOVAR (19 82) demonstrated its presence in the Egerian stage of Austria on basis of cuticular ana­lysis. The Kesztölc remains contain no epidermis, thus assignment of the species could be performed on morphological grounds only. The species, demonstrated by cuticular analysis from Miocene and Pliocene layers (KNOBLOCH and KVACEK 1976), can be considered a typical Neogene element, already present in the Egerian, where Palaeogeneous and Neogeneous elements occur together. Betulaceae Alnus L. Alnus sp. Material: MÁFI: BK-1123, BK-1124 The length of the measurable leaf is 4.4 cm, its width is 2. 3 cm. Its shape is obovate. The basis is acute, the apex is missing. The margin of the leaf is denticulated, the venation is craspedodromous . There are 6 pairs of secondary veins branching out from the midvein, which are slightly curved and terminate in teeth. The tertiary venation consist of a dense network of veins, perpendicular to the secondary veins. The specimen found slightly resembles the species A. julianaeformis (Sternberg) Kva­cek et Holy, but based on two specimens found altogether, as well as having no cuticle at all, they cannot be assigned unambiguously. The species of this genus belong to the Arctotertiary flora, some representatives, how­ever, like A. julianaeformis can persist in the subtropical climatic phases of the Miocene as well. The above species is a member of the swamp vegetation, which is not unlikely in the case of Kesztölc, in light of some other remains also coming possibly from marshy regions. The presence of the species Taxodium dubium denotes the presence of another type of vege­tation apart from the laurel-forest. Ulmaceae Ulmuls L. Ulmus pyramidalis Goepp. Pl. IX, Figs 2-3. 1845 Ulmus bronni Ung. partim; Unger, PI. 26, Fig. 1, non Figs 2-4. 1845 Ulmus longifolia Ung. ; ibid., Pl. 26, Figs 5, 6.

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