Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 92. (Budapest 2000)

Roháček, J. ; Papp, L.: Crumomyia microps sp. n. from Austria and notes on other cavernicolous Crumomyia species (Diptera: Sphaeroceridae)

Kahlen leg. (coll. Silesian Museum, Opava); 2 female paratypes, Austria: Osttirol, Karnische Alpen, Obstanser Eishöhle, 2180 m, 7.9.1997-15.9.1999, Kahlen leg. (coll. A. KOFLER and Silesian Museum, Opava). Etymology - The new subspecies is named according to large eyes (oculeus = big-eyed). Diagnosis - Male. Body length about 3.8^4.0 mm; general colour dark brown to black. Head with chaetotaxy and microtomentose pattern as described for C. parentela (cf. NORRBOM & KlM 1985). Gena relatively high but eye not reduced; its longest diameter 1.7-1.9 times as long as shortest genal height. Aristal cilia slightly longer than in C. p. alpicola. Mesopleuron with a narrow microtomentose band along dorsal margin, otherwise bare, thus pleura as in other taxa of the C. parentela complex. Thoracic spiracle normal, not reduced. Legs as in C. p. alpicola, normal in length and with similar chaetotaxy. Wing not reduced (length 3.7-3.8 mm, width 1.4-1.5 mm), with cross-veins slightly infuscated. Abdomen with sclerites of normal size. Male genitalia as in C. cavernicola cavernicola (see PAPP & ROHÁCEK 1983, Figs 12-14), thus with shorter, distally more dilated and rounded gonostylus, short hypandrial apodeme and anterior apical lobe of postgonite narrower than in C. parentela. Female. Postabdominal sclerites very similar to those of C. cavernicola cavernicola (cf. PAPP & ROHÁCEK 1983, Fig. 9); the relatively short and dorsomedially divided 8th sternum is the most diagnostic feature distinguishing female of C. cavernicola from the female of the sympatric (see above !) C. parentela alpicola. Discussion - The new subspecies is here recorded from both caves near Obstan. It is externally hardly distinguishable from C. parentela alpicola; however, it can be easily separated from the latter by the features in the genitalia. C. cavernicola, originally described as a subspecies of C. parentela, is elevated here to a species level because the above described C. cavernicola oculea ssp. n. occurs sympatrically with C. parentela alpicola in the Alps (also records of the latter from the Hohe Tauern Mts. are not far from the type locality of the new subspecies) and the comparison of their genitalia revealed distinct differences between these taxa. Nevertheless, both species (C. parentela and C. cavernicola) are very closely related, and have to be splitted not very long ago. In contrast to the nominate subspecies (C. c. cavernicola) the new subspecies C. c. oculea is morphologically unmodified despite the fact it was discovered in a cave, and, therefore, it cannot be treated as a troglobiont. It probably also live free in alpine zone of the Alps and can be classified at most as a troglophilous species, like C. parentela alpicola occuring besides mountains also in caves in Carpathians (Slovakia).