Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 85. (Budapest 1993)

Nesemann, H.: Identification key to the Hungarian leeches of the subfamily Trochetinae Pawlowski, 1954, with notes on systematics of the subfamily Erpobdellinae Blanchard, 1894 (Hirudinea)

SYSTEMATICS OF THE SUBORDER ERPOBDELLIFORMES SAWYER, 1986 Erpobdellidae BLANCHARD, 1894 Erpobdelliformes with more or less prominent pharyngeal pseudognaths of segment III. Northern Hemisphere. Twenty-five species are described from Europe, several of them are divided into valid subspecies, e.g. Dina lineata, D. absoloni and Trocheta subviridis. I. subfamily Erpobdellinae BLANCHARD, 1894 Genera Erpobdella DE BLAINVILLE, 1818 Nephelopsis VERRILL, 1872 Erpobdellidae with complete homonomous five-annulate segment, primary (a2) and secondary (bl, b2, b4, b5) annuli (Fig. 1), testisacs numerous, lacking pharyngeal stylets, gonopores on somite XII, caudal sucker always developed. Preatrial loop of male paired ducts in somite XI. Suctoroid oral sucker with small preoral segments I-II (Fig. 9). Northern Hemisphere: Holarctic (Erpobdella), Nearctic (Nephelopsis). Four species in Europe (see Checklist). II. subfamily Trochetinae PAWLOWSKI, 1954 Genera Dina R. BLANCHARD, 1892 Trocheta DUTROCHET, 1817 Archaeobdella GRIMM, 1876 Erpobdellidae with complete, heteronomous five- to nine-annulate segment, primary to quartary annuli, annulus b6 always widened and homonomously or heteronomously subdivided (Figs 3-4), testisacs numerous, lacking pharyngeal stylets, gonopores on somite XII or XIII, female gonopore sometimes shifted in caudal direction, caudal sucker sometimes reduced (Archaeobdella). Preatrial loop of male paired ducts extremely long, extended and not coiled atrial horns in somite XI. Rostelloid oral sucker with large preoral segments I-II, forming a typical rostellum (Figs 7-8). Palaearctis: Palaearctic (Dina), West Palaearctic (Trocheta), Ponto-Caspian Basin (Archaeobdella), several monophyletic endemic taxa of the Lake Ochrid (Dina). In the author's opinion, the status oiDina maoriana MASON, 1976 from New Zealand remains unclear. Twenty species with several subspecies are known in Europe (see Checklist), but a few more seem to exist (NEUBERT & NESEMANN, in print). III. subfamily Mooreobdellinae subfam. n. Genera Mooreobdella PAWLOWSKI, 1955 (type genus) Fadejewobdeila LUKIN, 1962 Erpobdellidae with complete heteronomous five- to six-annulate segment, primary to tertiary annuli, annulus b6 widened, lacking pharyngeal stylets, gonopores on somite XII, lacking preatrial loop of male paired ducts. Holarctis: Nearctic (Mooreobdella), Palaearctic Ponto-Caspian (Fadejewobdeila). One species in Europe (see Checklist).