Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 66. (Budapest 1974)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XII.

with a pale violet suffusion. Spiculae 7 mm long and 2,5 mm wide, 2—3-florous, puberulous or sparsely hirsute. Glume unequal, lower ones 5,4 mm long, narrowly lanceolate, upper ones 6 mm long, oval, acute or obtuse, lemmae 6 mm long, acute, puberulous, paleae 5 mm long, bicarinate, hyaline, acute. Antherae 2,5 mm long. Stomata 53 p. Though I have seen one plant only in the herbarium of the Botanical Institute and Botanical Garden of Zürich, there can be no doubt that this beautiful poly ploid grade species has a much wider range in the massif du Central — only it was misinterpreted. The plant appears under the name Koeleria cristata PERS. G. var. vestita (HÉRIB.) ROUY in ROUY'S work "Flore de France" (1913). The plant thrives also in my experimental garden, grown from a seed collected form a plant in its natural habitat and occurrence. The very habit of the plant reveals that it is a polyploid of Koeleria schroeteriana (DOM.) UJH. Both plants have narrow and viridescent basal leaves, pubescen t with thin macrohairs. The anatomical structure of the leaf epiderm is also qu

1te similar, with only the cells being bigger. The form of the panicles of the two species also resemble each other to a great degree : they are rather compact, cylindrical, with a violet suffusion, the spikelets hirsute, their form the same, with only that of Koeleria heribaudii UJH. being considerable larger. The similarities between the habits, morphological picture and the epidermal structure of Koeleria pyrenaica (DOM.)UJH., Koeleria britannica (DOM.)DRITCE var. pontariierii (DOM.)UJH., Koeleria schroeteriana (DOM.)UJH., and Koeleria heribaudii UJH. are not to be doubted. The slightly more removed Koeleria britannica (DOM.) DRUCE also shown a number of related features. These endemic species, inhabiting rather delimited and partly isolated localities, had broken away from each other during the glaciations or came into being in their réfugia, surviving there tens of thousands of years. The examination of the plants in their natural habitats, the study of live material grown in experimental gardens, and their genetico-evolutional investigation surely verify these statements. The problem K. DOMINAS of Subsectio Caespitosae verae Tribus Cristatae subtribus Fatisceutes The Latin diagnosis of this taxon is as follows: Vaginis vetustis in fila recta aut in lacinias angustissimas fatiscentibus. DOMIN assigned to this group two Asiatic species: Koeleria tokiensis DOMIN and Koeleria mongolica DOMIN, as well as two alpine species: Koeleria alpigena DOMIN and Koeleria Mannagettae DOMIN. The distance of many thousands of kilometres between the areas of two species each should in itself exclude the evolutional relationship of the four species mentioned above. As in several other cases, DOMIN constructed his Koeleria system on the basis of the application of fine life-form features. I have not seen the Asiatic specimens, but, on the basis of the examination of the types of the two European species, the subtribe is to be considered the result of an erroneous observation. The specimens of both Koeleria alpigena DOMIN and + Koeleria Mannagettae DOMIN are infested by mycelia of Fungi imperfecti. The infection is especially heavy on the type of Koeleria alpigena DOM., not only are the small tufted spots of the white mycelia observable among the remains of the vetust vagina and in the panicle, but even the conidia are present. There are less on the Koeleria Mannagettae