Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 66. (Budapest 1974)
Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XII.
A robust, loosely cespitose subalpine plant. Innovation extravaginal, therefore more loosely cespitose than Koeleria eriostachya PANC. Sward high, owing to the subalpine habitat conditions. Rhizome ramifying, owing to rich innovation frequently elongate recumbent; at other times rhizome compact, thick, the several shoots nearly adjacent to one another, the sward apparently similar to that of the intravaginal species. Rhizome 2—3 mm thick, occasionally even thicker by the covering remnants of root branches and vaginae, sublignosae. Plant generally 50 cm high, with some smaller and also taller specimens occuring. Vetust vaginae mostly 3 cm long, in drier and sunnier localities shorter, in shaded sites longer, glabrous, or very minutely sericeous, generally brownish-grey, sometimes lighter, or even rufescent. Senile blades rarely short, (8—10 cm long), generally 15 cm long, glabrous, explanate, or very weakly convolute; margins scabriusculous, green, underside pruinose greyish, auricle not conspicuous, apically attenuating, acuminate. Lowest blades slightly wider, 2—5 mm wide, higher ones narrower, with 2 cm long glabrous or glabriusculous and rufescent vaginae, (on some specimens from the Velebit with long, thin hairs). Ligulae very short, abrupt, minutely dentate. Juvenile leaves long, 20—25 cm, narrower, 1—1,5 mm wide, convolute, their undersides green, uppersides pruinose grey, marginally heavily scabriusculous, entirely glabrous, auriculate, auricle not conspicuous ; their vaginae generally 3 cm long, glabrous or sericeous owing to minute and appressed hair primordia. Ligulae somewhat larger, dentate, marginally finely dentate. Culm 30—40 cm long, 1 mm thick, below panicle densely hirsute. Specimens with a glabrescent culm appearing especially among exemplars from the Velebit ; their inflorescence is also generally glabrescent. Culm hirsute usually only below panicle, but in some plants antire culm hirsute. Panicle generally 5 cm long, in some specimens even 8 cm long (probably representing plants grown in better soil conditions. A specimen collected by E. KOVÁCS on the Triglav in 1958, at 1700 m a. s. 1., and having a panicle of 5—8 cm long, developed in my experimental garden 10—11 cm long, therefore pyramidate, panicles. On this plant also the innovational leaves attained a length of 35—40 cm). Panicles 1—1,2 cm thick, compact or slightly lobate, elongately cylindrical, or oval, apically rounded or (rarely) attenuating, rather basally interrupted, at flowering greenish violaceous, later slightly pale brownish, hirsute, or very frequently subhirsute. Spiculae generally 7 mm long. (I have repeatedly emphasized that, according to the ploidy grade, there is a certain intraspecific variability, depending partly on the given habitat conditions — a shallower or deeper humus content, water regime, etc. — and this holds also for Koeleria carniolica KERN.) Spiculae generally elongate, gracile, but also specimens with shorter and stouter spiculae occurring. (This difference, however, does not exceed 1 mm. The spiculae of the above mentioned specimens from the Triglav grew longer in the experimental garden, but these plants had optimum habitat conditions, — e. g. watered on the drier summer days). Glumae uneven, lower ones narrower, 4 mm long, upper ones 5 mm long, stouter, apically obtuse or slightly acute, rarely ciliate, lemmae 5 mm long, lanceolate, frequently attenuating aristately, sericeous with appressed pili, paleae 4 mm long, bicarinate, membranaceous, hyaline, acute. Antherae 3 mm long. Exemplars with wholly glabrous culms and strongly glabrescent or glabrous panicles are especially frequent among those collected at Sugarska-Duliba, Allan, Raduc, and Risnjak. Plants with pale yellowish or greenish panicles are also frequent among these. Their leaves are also glabrous, with pili present only occasionally on the margins of the lower part of the innovational blades. The basic structure of
