Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 66. (Budapest 1974)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XII.

the leaf epiderm is, however, similar to that of the typical specimens of Koeleria carniolica KEEN. There are slightly more silica-bodies in the costal zones, and since the leaves are often shorter, also the parenchymatous cells are shorter. As I had no occasion to visit the Velebit Range, and I have no plant from this area in my experimental garden, I cannot decide wheter these features came into being owing to the more xerothermous habitat conditions or if they are possibly identical with Koeleria carniolica KEEN. Some of these plants have been identified by DEGEN, LENGYEL, and even DOMIN as Koeleria montana (HAUSM.) D. T., or Koeleria splendens PEESL sensu DOMIN. They so not belong to the species Koeleria subcadata (ASCH. & GEAEBN.) UJH., or to Koeleria borbasii UJH., both relegated to the Koeleria splendens spec. coll. There is a possibility that they represent Koleria montana (HAUSM.) D. T. specimens, descending to the Velebit. In any case, I cannot give a definite anwer in this case at present, but, since they most resemble Koeleria carniolica KEEN, in habit and structure, I retained them in the list above. The exsiccata specimens of RIGO, HUTEE and POETA, originating from the Venica Alps, belong with hirsute or glabrescent panicles and hirsute culms to the typical Koeleria carniolica KEEN. It is only natural that I cannot now accept the views of either DOMIN or other authors that the types of diverse pubescence are transitional forms among various species. I merely refer here to my evolutional concepts discussed in my preceding paper (ÚJHELYI 1972), or my lecture read before the Symposium'on Evolution at Tihany, 1970. I have mentioned at the head of the chapter that I found also a quite gracile specimen with small panicle, definitely thin culm and leaves. Such is JABORNEGG'S plant: Koeleria carniolica KEEN. In rupibus calcareis montis Loibl Carinthiae, 1400 m. Aug. 1889. Scripta DEGENI

: non! Dr. Á. DE DEGEN. Schedula Domini: "Koeleria eristachya PANC. var. carniolica (KEEN.) DOM. Eigentlich eine Uebergangsform zu der K. montana. Die JL. carniolica scheint überhaupt ein Verbindungsglied bei der Arten zu sein. Besitze vom demselben Standorte Exemplare mit stärkere behaarten Spelzen. DOMIN". This is no circumstances a transitional form. Its spicule is expressedly slender, the glumae and lemmae are quite narrow and of a pale green hue. All these refer to the dimensions of a tetraploid grade plant. It resembles Koeleria carniolica KEEN, in having a hirsute culm under the panicle, itself is shinily glabrous. This rate of hirsuteness of the culm occurs also in Koeleria mollis MANN, Koeleria kernerii UJH., or in Koeleria montana (HAUSM.) D. T. The underside of the leaves of the two latter species is also glabrous, the margin — as in the plant indicated above — ciliate. It is possible that Koeleria montana (HAUSM.) D. T. occurs also in Carniolia, but it may also represent the progenitor of Koeleria, carniolica KEEN. I cannot decide this a single herbarial sheet alone. With reference to my previous papers, in which I have pointed out the wholly artificial specific notion of Koeleria pyramidata (LAM.) DC, and the existence of the evolutional series of the Series Ciliatae UJH., it is easily understandable that the complete clarification of taxa, nearly related as to lifeform and if very similar ecological requirements, is not so very simple when based on herbarial and not live material. Until very recent times, compilatory work is restricted to the usage of literature texts. In no other way could, e. g. BOBBÁS' notions from 1882, or HACKEL'S argumentations have influenced DOMIS'S ideas. Subsequent authors have simply taken over DOMIN'S expositions, and by misinterpreting these speculative assumptions some of DOMIN'S taxa have been retained, some degraded or straightaway synonymized — without ever having examined the plants themselves (HE-