Szekessy Vilmos (szerk.): A Magyar Természettudományi Múzeum évkönyve 62. (Budapest 1970)

Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria. IX.

The vetust vaginae are whole, occasionally irregularly lacerated. The rhizomes are short, hardly incrassate. The 3-4 cm long stolos are also gracile, narrow, about 0.7 mm thick. The senile leaves are comparatively short, at most 8 cm long, narrow, explanate, about 0.8 mm wide, glabrous, the very minute hairs only initial. When dry, they are slightly convolute, glaucescent, their vaginae at most 3 cm Jong, glabrous. The ligulae —in contrast with those of Koeleria javorkae UJH.,—are hardly developed, abrupt, marginally very finely crenate, the crenation apically subciliate. The juvenile leaves are elongate, 17-19 cm long, very narrow, in a dry state convolute, 0.3 mm wide, subglabrous, and owing to a very fine waxy coat pruinose. The vaginae of the juvenile blades are 4 cm long, with a hardly discernible auricle emitting, similarly to the situation in Koeleria javorkae UJH., 3-4 cilia of about 1.8 mm length. Culm in a wholly developed state foliose up to its half or occasionally slightly higher. The culm blades are 6-7 cm long and about 1.5 mm wide, explanate or slightly convolute, apically gradually attenuating, acute, their vaginae about 17 cm long. The ligulae are about 0.4 mm long, also abrupt and very minutely ciliolate. The culms are about 55 cm long, narrow, about 1-1.5 mm thick, glabrous. Similarly to the other two members of the Series, the panicle is cylindrical, compact, inferiorly interrupted, 6-11 cm long and about 7-10 mm thick, in the flowering state washed violet, later pale yellowish, shining or partly sericeous. The spikelets are 4-4.5 mm long and 1.5 mm wide, biflorous. The glumae are unequal in size, shiny, dorsally very minutely scabriusculous. The lower gluma is 4 mm long, very narrow, the upper one 4.5 mm long, apically acute. The lemmae are 4 mm long, narrow, apically acute, the paleae 3 mm long. The stomatal cells are 34 u. long. As evident from the description, the plant belongs to the relationship of the Series Javorkae UJH., while all measurements indicate that we have to do with a diploid grade member of the Series. That the species belongs to the Series mentioned above is substantiated by the presence of the stolos —unknown in any other Eureopean Series—the glabrescent leaves without pili, the convolute juvenile leaves lacking any protruding rib, the elongately cylindrical panicle and the epidermal structure on the abaxial side of the blades. In the members of other series, the parenchymal cells of the costal zone alternate only with silica-bodies, and if there appear occasionally cork cells — primarily in the innovational leaves of Koeleria eristata (L.) PEES. em. BORB.— they stand invariably alone, that is, joining the parenchymal cells on both sides. They are general only in the members of the Series Javorkae UJH. In addition, the cork cells join the parenchymal cells always as a twin cell of a silica body. It is readily understandable that Dr. E. GY. NYÁRÁDY, Academician, identified his specimens as Koeleria gracilis PERS. glabra JANKA. From Koeleria javorkae UJH., the new species differs primarily by its external habits and more gracile stature. Koeleria javorkae UJH. is a tetraploid grade plant, whereas Koeleria nyaradyi UJH. is doubtless a diploid as attested by all its measurements. Koeleria nyaradyi UJH. inhabits the oakwood clearings of the Transsylvanián Basin, at 550-600 m a.s.l. On the other hand, Koeleria javorkae UJH. is a plant of I he lussocky moors and peats of the Great Hungarian Plains, ascending the hillsides only along the eastern borders. The structure of the leaf epiderm shows the most characteristic contrast against that of Koeleria javorkae UJH. Whereas on the abaxial side of the innovational leaves of this latter species the parenchymal cells are large in the intercostal zone, they are minute in Koeleria nyaradyi UJH., Iii TT. Múzeum JÉvkünyvu 1070