S. Mahunka szerk.: Folia Entomologica Hungarica 63. (Budapest, 2002)

nation. 77z. celelata is a supralimital member of Micandra; its female has female genitalia qualitatively identical with Micandra species and wings in both sexes showing the typical Micandra venation (Röber 1892: 265, Pl. 146, fig. "Micandra Schatz Platyptera Feld."), the latter being stressed as an important character by Robbins (1986: 154); I therefore transfer the species from Thecla to Micandra, thus, Micandra celelata (Hewitson, 1874), new combination. Placement in Laothus - Besides Thecla barajo Reakirt, 1867 and Thecla laothoe Godman et Salvin, 1887, Johnson et al. (1997: 23) placed Pseudolycaena danaus in Laothus Johnson, Kruse et Kroenlein, 1997 (type species: Thecla ba­rajo, by original designation) but they remarked that the species composition of the genus was based primarily on wing pattern and morphological characters. I have dissected female specimens of the type species of Laothus along with superficially similar taxa Pseudolycaena viridicans Felder et Felder, 1865, Thecla erybathis Hewitson, 1867 and Thecla oceia Godman et Salvin, 1887, and discovered that they possess almost identical female and male genital structures and differ from Margaritheclus; females of the former species have the female ductus bursae sclerotized equally everywhere along the tube; males lack the lateral projection of the valva. In addition, they also differ strongly in wingshape, colouration, and ven­tral pattern. Additionally, Laothus and Gibbossa Salazar, 2001 (replacement name of the homonym Gibbonota Salazar et Lopez, 1996, type species: Thecla gib­berosa Hewitson, 1867, by original designation, cf. Salazar 2001) now appear to be monophyletic, if not synonymous, as male and female genital structures, along with ventral wing markings of these "genera" are qualitatively not distinct. The type species of the two genera represent two extremes within a uniform mono­phyletic group since the taxa erybathis and phydela are intermediate, filling the morphological gap between barajo and gibberosa. This monophyly has already been expressed by Luis-Martinez et al. (1995: 127) as well as by d'Abrera (2001: 195) who placed all the taxa mentioned above under "Gibbonota". His action was actually based on my studies (cf. d'Abrera 2001: 194) which is why evidence was not given by d'Abrera supporting such synonymy. Accordingly, I here formalize all these taxonomic changes and place Laothus and Gibbossa as synonyms; there­fore Gibbossa Salazar, 2001 = Laothus Johnson, Kruse et Kroenlein, 1997, new synonym, thus, Gibbossa viridicans (Felder et Felder, 1865), new combination, Gibbossa erybathis (Hewitson, 1867), new combination, and Gibossa barajo (Reakirt, 1867), new combination and Gibbossa oceia (Godman et Salvin, 1887) new combination. Relationships of Margaritheclus - Wingshape of Margaritheclus amongst eumaeine lycaenids is not peculiar. The male androconial cluster, what I suspect is an androconial patch, indicates a closer relationship with Areas Swainson, 1832

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