S. Mahunka szerk.: Folia Entomologica Hungarica 63. (Budapest, 2002)

(type species: Papilio imperialis Cramer, 1775, by monotypy) and Paiwarria Kaye, 1904 (type species: Papilio venulius Cramer, 1779, by original designation); however, wing pattern and genital characters do not support this context. The ventral wing pattern of Margaritheclus is also not unique qualitatively. There are several genera with vertical patterns of ruptive gleaming lines or streaks reminiscent, or nearly identical with, Margaritheclus. These include Salazaria d'Abrera et Bálint, 2001 (type species: Thecla sala Hewitson, 1867, by original designation), Timaeta Johnson, Kruse et Kroenlein, 1997: 23 (type species; Pseudolycaena timaeus Felder et Felder, 1865, by original designation) and Janthecla Venables et Robbins, 1991 (type species: Thecla janthina Hewitson, 1867, by original designation). Characters of male androconia do not support the relationship of Timaeta and Janthecla but Salazaria, which lacks male dorsal androconia, is closer to Margaritheclus, a situation also supported by the female genitalia. All of these genera possess a more or less sclerotized posterior lamella in the female. However, it is difficult to judge at the current state of knowledge concerning eumaeine lycaenids, whether the similarities in these structures indicate monophyly or homoplasy. The male genital configurations of Margaritheclus are typical eumaeine with a thin tegumen, loose manica, thin and long penis, and long saccus typical of a cluster of genera (cf. Robbins 1986: 153). Morever, Margaritheclus valvae possess a laterally occurring, posteriorly pointed process which is almost as long as the genital's saccus. This process seems to be unique amongst eumaeines. This male genital configuration, associated with a long female ductus bursae, simple cervix, and large posterior lamella also apparent in Margaritheclus may well suggest that Margaritheclus is one of the most conspicuous eumaeine lycaenid genera. SUMMARY I have demonstrated that the original combination, and all subsequent placements of Pseudolycaena danaus Felder et Felder, 1865, were erroneous from a phylogenetic point of view. All taxa previously clustered with the danaus species have to be placed in other genera, resulting in various new combinations (in alphabetical order): Atlides havila, Denivia adamsi, Gibbossa erybathis, G. oceia, G. viridicans and Micandra celelata along with the synonymization of the genus Laothus with Gibbossa. I have erected the genus Margaritheclus with the type species Pseudolycaena danaus and also described the sister species of P. danaus, each differing in characters of the ventral wing pattern, male dorsal androconial cluster, and female genital structures. In addition I have described another species, M. dabrerus (whose extant material lacked abdomen) placing it in Margaritheclus on