S. Mahunka szerk.: Folia Entomologica Hungarica 60. (Budapest, 1999)

in PBR were brachypterous. Because of this fact it is probable that there are habitats within the given sites, where they maintain stable populations in which dispersal is not necessary (Liebherr 1986), similarly to the above listed most common species. A large part of the relatively small distribution area of C. scheidleri is found in Hungary. Its preferred biotopes are the following: Arrhenatheretea, Querco-Fagetea and agricultural land. Pavlicek and Houskova (1989) listed fields, meadows and garden as characteristic habitats for C. scheidleri. Kromp (1990) recorded numerous specimens in Austrian potato fields. C. scheidleri in higher number has been shown in beech forests by Korbel (1973), and in hedges, watersides and various types of forest. P. melas was abundant in mixed oak forests of PBR according to data of Báldi and Ádám (1991) and Fazekas et al. (1992) whose assessed their results by pitfall trapping in the Buda Hills near Budapest, Hungary. The formerly five species represented about 60 per cent of all individuals collected. Thus, they probably are important components of the Coleoptera fauna in PBR because of their high number of individuals. Of the rest, 13 more frequent species (Table 1)12 are also forest carabids (Thiele 1977), and as generally known are hygrophilous, dark prefering and nocturnal in habitat (Thiele 1969). One species, Harpalus rufipes is a true 'ruderal' ground beetle and it is a typical species of open areas, especially agricultural fields (Luff 1987; Lövei and Sárospataki 1990). Among the carabids of moderate frequency (< 1%) several species have high affinity to forest, including e.g. Harpalus marginellus, Licinus hoffmannseggi and Trechus pilisensis. They are montane species and usually occur in beech forests and/or of those associations. On T. pilisensis there is no information, there are only distribution data, as it was given by Korbel (1973), Sustek and Zuffa (1987), and in Hungary e.g. Csiki (1946); Horvatovich (1991) and Szél (1996). The 102 adults of T. pilisensis were brachypterous. H. marginellus is a brachypterous but polymorphic species (Brandmayr and Zetto Brandmayr 1988) and spring breeder with an inclination to phytophagous behaviour (Brandmayr 1974). Distribution data were given by former authors and lately Fig. 1. The 15 sites in the Pilis Biosphere Reserve, Hungary, clustered according to the distances of their carabid faunas (data from 1982-84)