S. Mahunka szerk.: Folia Entomologica Hungarica 60. (Budapest, 1999)

by Horvatovich (1991) and Szél (1996). L. hoffmannseggi is well enough known. Key to determine its helicophagous larvae is given by Arndt (1991). The imago is brachypterous (Brandmayr 1983) and autumn breeder (Brandmayr 1974). L. hoffmannseggi is a highly specialised oligophagous predator carabid species. Feeding behaviour pattern of imagoes (so called "shell-crushing") is supported by its enlarged flat head, asymmetric mandibles and labrum, and the broadly excavated clypeus (Brandmayr and Zetto Brandmayr 1986). The molluscivory is a general feature of licinines (Ball 1992). Generally, specimens of L. hoffmannseggi were caught in Fageium association of Fagetum, e.g. in Romania by Varvara et al. (1993), in Slovenija by Brandmayr (1974, 1983), in Switzerland by Borcard (1982). In Croatia, L. hoffmannseggi is a characteristic species of Abieti-Fagetum illyricum (Durbesic 1984). An ecobiogeographical characterisation of this ground beetle is given by Brandmayr and Zetto Brandmayr (1988). Our traps caught 23 imagoes on six sites of PBR. Thus, based on our results we may conclude that L. hoffmannseggi is more widespread in Hungary than it was found by Tóth (1980) earlier. This statement is supported by the data of Magura et al. (1997). The results of the hierarchical clustering and PCA ordination of 15 sites based on their carabids data are presented in Figs 1-2. It can be seen from the dendrogram that the clusters form four groups: A, B, C and D. These groups are separated from each other primarily along axis one of the ordination graph. Groups are described as follows: Group A: it is a discrete group dominated by H. rufipes and Carabus ullrichi, but some species occurred with moderate frequency, including Platynus dorsalis and Brachinus explodens. Group B: also it is a discrete group, characterized by species such as F melanarius. A colouring element is Carabus violaceus here. Fig. 2. PCA ordination plots, axis 1 by axis 3, based on their ground beetle species data. Plots joined to ellipses according to their cluster groupings (A-D) obtained from the cluster analysis (see Fig. 1; the given numbers represent the sampling sites)