S. Mahunka szerk.: Folia Entomologica Hungarica 60. (Budapest, 1999)

Similarly to the former species, P. melanarius is also an "exotic" ground beetle (Niemelä and Spence 1991), but contrary to A. parallelepipedus it is very eurytopic (Turin and Heijerman 1988). P. melanarius specimens can occur in relative high numbers in quite different habitat types which do not have connection to forest. For example, it is a common species of the agricultural landscape (cf. Thiele 1977), where it is considered an important biocontrol agent of aphids and other arthropod pests (e.g. Hance et al. 1990). P. melanarius is a dimorphic species (Lindroth 1945), but all individuals captured Table 1. List of ground beetle species caught in 15 topographically different trapping sites of the Pilis Biosphere Reserve, Hungary, in 1982-84 Mean rel. No. of Total Species frequency occupied sites Aptinus bombarda (Illiger, 1800) 23.43 12 4156 Abaxparallelepipedus (Piller et Mitterpacher, 1783) 15.07 15 2250 Carabus scheidleri Panzer, 1799 7.97 14 750 Pterostichus melas (Creutzer, 1799) 7.94 15 635 Pterostichus melanarius (Illiger, 1798) 5.57 12 693 Carabus convexus Fabricius, 1775 4.60 10 121 Carabus nemoralis O. F. Müller, 1764 3.37 13 296 Pterostichus oblongopunctatus (Fabricius, 1787) 3.26 14 520 Carabus ullrichi Germar, 1824 3.25 10 225 Carabus coriaceus Linnaeus, 1758 2.52 14 217 Harpalus rufipes (De Geer, 1774) 2.12 9 92 Harpalus atratus Latreille, 1804 2.08 8 119 Platyderus rufus (Duftschmid, 1812) 1.85 9 182 Carabus hortensis Linnaeus, 1758 1.81 9 265 A bax ovalis (Duftschmid, 1812) 1.55 10 286 Abaxparallelus (Duftschmid, 1812) 1.23 9 214 Molops piceus (Panzer, 1793) 1.21 12 162 Platynus assimilis (Paykull, 1790) 1.20 6 194 Further species: Platynus dorsalis (Pontoppidan, 1763), Brachinus explodens Duftschmid, 1812, Carabus violaceus Linnaeus, 1758, Cychrus caraboides (Linnaeus, 1758), Laemostenus terricola (Herbst, 1784), Trechus pilisensis Csiki, 1918, Amara convexior Stephens, 1828, Notiophilus rufipes Curtis, 1829, Pterostichus niger (Schaller, 1783), Abax carinatus (Duftschmid, 1812), Epaphius secalis (Paykull, 1790), Bembidion guttula (Fabricius, 1792), Leistus rufomarginatus (Duftschmid, 1812), Calosoma inquisitor (Linnaeus, 1758), Harpalus marginellus Dejean, 1828, Amara saphyrea Dejean, 1828, Pterostichus ovoideus (Sturm, 1824), Poecilus versicolor (Sturm, 1824), Stomis pumicatus (Panzer, 1796), Pterostichus strenuus (Panzer, 1797), Calathus fuscipes (Goeze, 1777), Brachinus crepitans (Linnaeus, 1758), Ophonus nitidulus Stephens, 1828, Licinus hoffmannseggi (Panzer, 1797), Harpalus tardus (Panzer, 1797), Harpalus rubripes (Duftschmid, 1812), Calathus cinctus Motschulsky, 1850, Callistus lunatus (Fabricius, 1775), Dromius spilotus (Illiger, 1798), Amara familiáris (Duftschmid 1812), Synuchus vivalis (Illiger, 1798), Nebria brevicollis (Fabricius, 1792), Amara apricaria (Paykull, 1790), Dyschirius globosus (Herbst, 1784), Poecilus cupreus (Linnaeus, 1758), Carabus intricatus Linnaeus, 1761, Amara ovata (Fabricius, 1792), Amara similata (Gyllenhal, 1810), Carabus cancellatus Illiger, 1798, Dromius quadrimaculatus (Linnaeus, 1758), Ophonus azureus (Fabricius, 1775), Amara aenea (De Geer, 1774), Harpalus tenebrosus Dejean, 1829, Bembidion deletum Audinet-Serville, 1821, Cymindis axillaris (Fabricius, 1794), Trechus quadristriatus (Schrank, 1781), Amara consularis (Duftschmid, 1812), Bembidion lampros (Herbst, 1784).