B. Papp szerk.: Studia Botanica Hungarica 39. 2008 (Budapest, 2008)

Erzberger, P.; Schröder, W.: The genus Schistidium (Grimmiaceae, Musci) in Hungary

Abaxial side of costa with scattered low papillae in upper part; leaf margins papillosedenticulate in upper part, more rarely smooth, broadly and strongly recurved throughout or in upper 3/4-4/5 of leaf length, ± bistratose. Lamina with scattered dorsal papillae, particularly near costa, and mostly with few ventral papillae, unistratose with bistratose spots and striae in upper part. Lamina cells in upper part rounded, slightly sinuose, 8-11 urn wide, in lower part wider and sinuose to nodulose, basal marginal cells with slightly thickened crosswalls. K+ red. Sporophytes abundant, shallowly immersed. Urn red-orange-brown, glossy, length/ width ratio (1.3-)1.5-1.9-2.3(-2.5) (BLOM 1996), in Hungarian material 1.78-1.83, n = 2. Exothecial cells predominantly isodiametric and transversely rectangular, but also rectangular, forming an irregular pattern. Stomata 4-9 per urn. Peristome teeth 310-420 urn, squarrose-recurved, red, gradually tapering to a fine point, from entire to strongly and irregularly perforated in central part, densely papillose. In Hungary, 5. confusum is the only member of the Strictum subgroup that grows on calcareous substrates, whereas S. papillosum and S. pruinosum grow on siliceous rocks. From those species, S. confusum differs by its less papillose lamina (Fig. 1 IE), with more papillae on the abaxial, fewer on the adaxial side of the leaf (S. pruinosum is strongly papillose on both sides: Fig. 30G, H). S. papillosum is similar in this respect (usually more papillae on the abaxial side than on the adaxial side), but differs by the lack of a distinct central strand and the peristome teeth erecto-patent (Fig. 25G, but squarrose when old). Other differences between S. confusum and 5. pruinosum include the wider and more sinuose lamina cells (Figs 1 ID, 30F) and the narrower perichaetial leaves of the former species (Figs 1 IC, 30D). Habitat: In Hungary, S. confusum was found on exposed concrete adjacent to dolomite. In Scandinavia, it grows in similar situations, on exposed calcareous substrates (BLOM 1996). Associated bryophytes: None. Vertical distribution: 250 m a.s.l. Fig. 12. Distribution of Scbistidium confusum