B. Papp szerk.: Studia Botanica Hungarica 32. 2001 (Budapest, 2001)

Hably, Lilla: Rhamnus warthae Heer, an important element of the Central Paratethyan region during the Egerian

mate of its habitat should be taken also into consideration. One of such possible events was the major sea level drop at 30 Ma (HAQ 1991). Otogenic phase at the boundary of early and late Oligocène also caused a change of the land surface, which must have had an effect on the ecology and environment of the flora. Connected to this floral change several new species appeared in this region, and were limited to the Egerian or Egerian-Eggenburgian ages, among them Alnus oligocaenica Andr., Ulmus pseudopyramidalis Z. Kvacek et Hably, "Talauma" egerensis Andr., Tetracentron agriense (Andr.) Z. Kvacek et Hably, Acer tricuspidatum Bronn f. hungaricum (Andr.) Z. Kvacek et Hably, Dryophyllum callicomifolium (Andr.) Z. Kvacek et Hably, "Arbutus" preunedo Andr., Ilex? andreanszkyi Z. Kvacek et Hably and Tuzsonia hungarica Andr. These species are know only from Hungary, and partly from Romania (Transylvania, Zsil Valley) from Egerian layers. It confirms, that the above-mentioned floras must have developed in close connection with each other, but isolated from other regions. We have to consider the presence of Magura ocean (CSONTOS et al. 1992) to the north, which might have formed a large barrier between the Hungarian and other floras located to the north. To explain the isolation of the Egerian floras in the Intra-Carpathian region, we also have to consider the palaeogeographical conditions according to the model of terrains (CSONTOS et al. 1992, HABLY and KÁZMÉR 1996). Acknowledgement - This work was supported by the Hungarian Scientific Research Fund (OTKAT 029041). REFERENCES ANDREÁNSZKY, G. (1966): On the Upper Oligocène Flora of Hungary. Analysis at the site at the Wind Brickyard, Eger. - Studia biol. hung. 5: 1-151. CSONTOS, L., NAGYMAROSY, A., HORVÁTH, F. and KOVAC, M. (1992): Tertiary evolution of the Intra-Carpathian area: a model. - Tectonophysics 208: 221-241. ERDEI, B. and WILDE, V.: An Oligocène leaf mac roflora from Csolnok (Dorog basin, N Hungary, (in progress) GlVULESCU, R. (1973): Ein Beitrag zur Kenntnis der fossilen Flora des Schiltals. - Mémoires 19: 7-27. HABLY, L. (1988): Egerian fossil flora from Kesztölc, NW Hungary. - Studia bot. hung. 20: 33-61. HABLY, L. (1989): The Oligocène flora of Nagysáp. - Fragmenta min. et pal. 14: 83-99. HABLY, L. (1990): Egerian plant fossils from Vértesszőlős, NW Hungary. - Studia bot. hung. 22: 3-78. HABLY, L. (1993a): Egerian macroflora from the Andornaktálya sandpit (Hungary). - Studia bot. hung. 24: 5-22.