L. Hably szerk.: Studia Botanica Hungarica 22. 1990 (Budapest, 1990)

Material: No.: 76.3.1.; 76.5.1.; 76.6.1.(=76.188.1. counterpart ) ; 76.7.1.; 76.11.1.; 76.14.1.; 76.21.1.; 76.101.1.; 76.107.1.; 76.130.1.(2); 76.136.1.; 76.141.1.; 76.153.1.; 76. 160.1.; 76.161.1.; 76.188.1.; 76.194.1.; 76.195.1.(2); 76.223.1.; 76.229.1.; 76.269.1.; 76.272.1.(4); 76.283.1.; 76.291.1.; 76.295.1.(2); 76.296.1.; 76.301.1.; 76.306.1.; 76.310.1.; 76.311.1.; 76.312.1.; 76.315.1. 37 pieces Description: The leaves are large, found generally without the petiole. Their shape is obovate, reaching their maximal width at the upper third of the lamina. This is 6.3 cm in case of the biggest piece, ranging generally between 3.0-5.0 cm. The complete length of the biggest leaf (76.6.1) comple­mented could be 16 cm, the total length measurable is 15.5 cm due to the fragmented state of the apex. The completed length of the rest of the leaves is ranging between 10-11 cm. The margin of the leaf is entire, the apex and the basis are acute. The biggest specimen is almost complete, thus the system Df the veins can be studied very well. At 1.2 cm from the end of the lamina, there are two strong basal veins branching off asymmetrically from the midve­in, having the same width as the midvein itself. The starting point of these veins is not always the same, they are generally starting with 0.2-0.5 cm of difference. In their distance from the basis there are also smaller differen­ces encountered, roughly proportional to the size of the leaf. On the specimen 76.6.1., the two basal veins are 1.4 cm apart from the midvein and 1.7 cm far from the margin at the widest point of the lamina. In case of some other specimens, however, they are nearer the margin, and in other cases, the distance from the margin and the midvein is roughly equal here . .Around the upper third of the leaf, there is a stronger secondary vein starting from the midvein, which is turning back towards the midvein at a higher point, forming a loop. The basal veins of second order contact this secondary vein, also in a loop-like manner. At the lower two-third of the leaf, veins of the third order are running between the midvein and the basal veins almost horizontally, branching and later anastomizating . At the side facing the margin there is a system of vigorous veins starting from the basal veins, taking an arched course and returning at some distance to the starting vein, creating a more or less regular loop system in the meantime. Apart from the Vértesszőlös site, the species was found on several Upper Oligocène palaeobotanical localities of Hungary (HABLY 1982, 1968). Close analogies of the species were found in the Zsil-vallsy ( Transylvania ) in the form of prints, published by STAUB ( 1887 ) under the name of Cinnamomum poly- morphum Al. Br. More new finds were published by KVAÖEK and WALT HER ( 1974) from the localities Habichtswald, Meissner, Seifhennersdorf . In the Vértes­szőlős flora, it is occurring associated with Palaeotropical and Arctotertia­ry elements, the composition of which is related in many ways to the Seifhen­nersdorf and the Zsil-valley floras. According to the taxonomical system of KVACEK and WALTHER (1974), the species D. cinnamomifolia is characteristic of the boundary of the Oligocène and the Miocene periods. Considering the fact that our locality is dated to the Egerian stage, this statement is corroborated by the Vértesszőlös evidence as well. The au­thors of the cited paper consider these large leaves as mesomorphic shadow leaves, while that of the D. lanceolata Ung. which is smaller and more narrow would be the xeromorphic light-lsaf. In the Vértesszőlös flora, both species (or types) are occurring, that is, not excluding that the remains belong to the same species as shadow- and light-leaves, respectively. From the point of climatology, the D. cinnamomifolia can be regarded as authoritative. In accord with the rest of the plants preferring humid envi­ronment we can state that this plant was the constituent of a subtropical forest abounding in humidity, growing D. lanceolata type leaves in the outer parts of the foliage exposed to light. Ii'

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