Dr. Murai Éva szerk.: Parasitologia Hungarica 20. (Budapest, 1987)
with thick walls, provided with long retractor muscle directed anteriorly to the middle of anterior margin of proglottis. Evaginated cirrus not observed but armament seen in cirrus duct consisting of triangular spines about 0.002 long. Vagina opening dorsally to male orifice; near its orifice armed with small spines; copulatory part 0.025-0.030 (av. 0.028, n=6) wide, conductive part 0.007-0.010 (av. 0.008, n=6) wide. Seminal receptacle developed in last mature, postmature and gravid proglottides; situated near vitellarium; elongate, with measurements 0.083-0.120x0.033-0.055 (av. O.lOlx 0.044, n=10). Vitellarium situated medially, slightly lobate. Ovary with two wings, aporal wing slightly larger than poral; aporal wing consisting of 8-13 lobes, poral one of 6-9 lobes. Ovary occupies anterior half of median field. Mehlis' gland distinct. Uterus in early stages reticular; with development of the eggs, it gradually occupies almost the whole proglottis. Eggs oval or subov^l. Embryonal hooks three pairs, with length 0.0140.016 (av. 0.015, n=15). Remarks The present results are in good agreement with the original description of this species (KOTLÁN, 1923). According to MATHEVOSSIAN (1963), on the basis of the similar number and shape of rostellar hooks, the number of testes and position of genitalia, M..^horvathi must be regarded a synonym of M. leptophallus. This opinion has been accepted by SPASSKY (19 65) but in the later book of the same author (SPASSKAJA and SPASSKY, 1977) M. horvathi is considered a valid species. Our results also confirm the validity of M. horvathi. It differs from M. lep tophallus in (1) the shape of the rostellar hooks, particularly those of the anterior row; (2) the length of the cirrus sac (0.320-0.475 in M. leptophallus and 0.148-0.210 in M. horvathi) ; and (3) the different structure of the vagina, especially the presence of armament in the copulatory vagina of MJioj-vjithL Nevertheless, all the other anatomical details, such as the presence of the retractor muscle of the cirrus sac, the position of the vagina dorsal to the cirrus sac, structure of the scolex, position of the genital glands, etc., which are very similar in both species discussed, suggest that they are closely related forms belonging to a monophyletic group. The genus Megacirrus Beck, 1951 has been erected for M. megapodii Beck, 1951 from a scrub hen species Megapodius laperouse senex in Palau Island (South Pacific) (BECK, 1951). The morphological features of this species are very close to those of M. leptophallus and M. horvathi. These three species are characterized by a similar structure of the Figs 24-29: Kotlanolepis yorkei (Kotlan, 1923) comb. n. (24= Scolex, scale bar 0.25; 25-26= Anterior and posterior rostellar hooks, scale bar 0.05; 27= Mature proglottides; 28= Postmature proglottides; 29= Gravid proglottides). Figs 30-34: Megacirrus leptophallus (Kotlán, 1923) (30= Eggs in Berlese solution, scale bar 0.03; 31= Postmature proglottides; 32= Rostellum with hook crown, scale bar 0.10; 33-34= Anterior and posterior rostellar hooks, scale bar 0.05). Figs 35-38: Megacirrus horvathi (Kotlán, 1923) (35-36= Anterior and posterior hooks, scale bar 0.05; 37= Rostellar hooks in various position, scale bar 0. 1; 38= Mature proglottis). (Photographs by Boyko B. GEORGIEV, Nos 25, 26, 33, 34, 35, 36 by Béla LOVAS)
