Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

64 external sclerites (possibly remnant of sternum X), ring structure: 0: present, well-developed, attached to abdominal segments (Figs 82-83), 1 : present, a (more or less) small ring, free from segments (Figs 76-79, 80-81), 2: absent. Female genitalia 65 spermatheca: 0: sac-like (usually highly unsclerotized) (Figs 98-99), 1: characteristically shaped, sclerotized (Figs 100-105). 66 receptacle: 0: undivided (Fig. 100), 1: divided (Figs 101-105). 67 spermathecal gland: 0: visible (Figs 99, 101, 103), 1: invisible (Figs 98, 100). 68 introversion on spermatheca (tubular portion penetrating distal bulb): 0: absent, 1: present (Figs 101-102). 69 umbilicus on spermatheca: 0: absent, 1: present (Figs 104-105). 70 sclerotized "tube" apparently associated with female genital tract: 0: absent, 1 : present (Fig. 84). Results The strict consensus tree (L=266, CI=36, RI=64) is shown on Fig. 2. Character states were distributed in the tree using Acctran optimization (favors reversals). For such an ancient group, the frequency of reversals might actually be high, therefore this optimization was favored against Deltran. As for the tribal classification of Oxytelinae, the following conclusions can be drawn: The tribe Deleasterini is likely not monophyletic, but as the basal clade of the subfamily with obviously great disparity, it makes sense to maintain it, pending further phylogenetic analysis. The tribe includes relict genera Milosynum, Oxypius and Euphemias. None of the included 6 genera exceeds 10 species, and 3 of them monobasic. The clade is apparently very ancient and has suffered a lot of extinctions. The tribe Coprophilini is sampled by a single taxon. The tribe Planeustomini is revalidated herein to include genera Planeustomus and Manda. The clade is supported by 3 unique synapomorphies: Clypeus reduced to narrow strip (1-1), eyes extending on ventral portion of head (3-1), mandibular mola much reduced or absent (6-1). The monophyly of this group is very likely. There is some difference in opinion on whether the name Planeustomini published originally in non-latinized form should be regarded available from JACQUELIN DU VAL, but since at this time it cannot be said for sure exactly how many times this extremely old name can be found (there is at least one in ÁDÁM and HEGYESSY 2001), it is considered the valid name here.