Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

species groups possessing peculiar character states or combination of character states are represented. The proper representation of the diversity of the ingroups required inclusion of five species, which have not been previously named. With one exception these taxa were discovered in the course of revisionary work and will be included in the publications that revise these genera or species groups. However, in order to properly associate these unique taxa with an identity, the descriptions are lifted and published here as an appendix (Appendix B) to validate the names. Character set and coding The selection of the characters was done from the perspective of the phylogenetic information content of features in a higher level classification. Characters that may be useful in species-level identification provide no phylogenetic resolution for higher taxa. The phylogenetic information in characters might vary greatly between genera. HERMAN'S (1970) character set was used as a basis, but many of his characters were either irrelevant for this analysis or proved to be highly variable and difficult to code. Therefore about two-thirds of his characters were modified, redefined or substituted by suites of characters almost completely omitted previously. The most important of these are the characters in the mouthparts, terminalia and genitalia. Characters and character states used to infer the phylogenetic relationships of the Carpelimus group (Oxytelinae) are as follows. Head 01 clypeus: 0: well developed, more or less rectangular (Fig. 25), 1: reduced to narrow strip (Fig. 26). 02 epistomal suture: 0: present, 1: absent. 03 eyes: 0: not extending onto ventral portion of head (Fig. 25), 1: extending onto ventral portion of head (Fig. 26). 04 gular sulci: 0: entirely separated (although anterior portion may run very close) (Figs 26-27), 1 : confluent at anterior portion (Fig. 25), 2: confluent in posterior portions, not anteriorly (Fig. 28). 05 head dorsal midline keel: 0: absent, 1: present. 06 mandibular mola: 0: separated, well-developed (Fig. 29), 1: much reduced or absent (Figs 30-31). 07 labrum at mid-line: 0: undivided (Figs 24, 32-33, 35), 1: midlongitudinally divided (except perhaps at very base) (Fig. 34).