Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Makranczy, Gy.: Systematics and phylogenetic relationships of the genera in the Carpelimus group (Coleoptera: Staphylinidae: Oxytelinae)

later. Consequently, the lineages found there must have originated earlier than the separation; if true, then some of the genera now recognized were already differentiated prior to 100 Mya. One can argue that the arrival of some groups might have happened later than the separation of New Zealand from Gondwana; the chance for this depends on a lot of poorly studied factors, like oceanic currents and the dispersal ability of oxyteline species. However, such long-distance, Overwater dispersal does not appear to be common, for example the whole subfamily Steninae, members of which occupy rather similar habitats to those of Oxytelinae, is absent from New Zealand (KLIMASZEWSKI et al. 1996). MATERIALS AND METHODS Material studied Out of the ten genera of the ingroup, the study of Ochthephilus, Xerophygus and Trogactus was done with a revisionary approach. The rest of the genera were investigated also on a worldwide basis, but the selection of taxa for detailed study was focused on faithfully representing their known diversity in the analysis. The creation of species groups based on real phylogenetic affinities was a goal, but for the larger genera (Carpelimus, Thinodromus) such systems are still preliminary and incomplete and subject to many changes as more taxa are described or studied. For this project, nearly ten thousand specimens were studied, either as part of the revisions or material necessary to assess the variability of characters in other genera. Although specimens for this project was borrowed from a much greater number of institutions, the following list is confined to the depositories of the material actually studied for the analysis: AMNH = American Museum of Natural History, New York City, New York, USA; BMNH = The Natural History Museum, London, England; CASC = California Academy of Sciences, San Francisco, California, USA; FMNH = Field Museum of Natural History, Chicago, Illinois, USA; HNHM = Hungarian Natural History Museum, Budapest, Hungary; ISNB = Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium; MHNG = Muséum d'Histoire Naturelle, Geneve, Switzerland; MNNC = Museo Nációnál de História Natural, Santiago, Chile; MRAC = Musée royal de l'Afrique centrale, Tervurcn, Belgium; MZLU = Museum of Zoology, Lund University, Lund, Sweden; NHMB = Naturhistorisches Museum, Basel, Switzerland; NHMW = Naturhistorisches Museum Wien, Vienna, Austria; NHRS = Naturhistoriska Riksmuseet, Stockholm, Sweden; NMPC = National Museum (Natural History), Prague, Czech Republic; NZAC = New Zealand Arthropod Collection, Landcare Research, Auckland, New Zealand; SEMC = Natural History Museum, University of Kansas, Lawrence, Kansas, USA; SMNS = Staatliches Museum für Naturkunde in Stuttgart, Stuttgart, Germany; USNM = National Museum of Natural History, [formerly: United States National Museum], Washington D.C., USA; ZMHB = Museum für Naturkunde der Humboldt Universität, Berlin, Germany; cSch = private collection of MICHAEL SCHÜLKE, Berlin, Germany; cGdR = private collection of GUILLAUME DE ROUGEMONT, Mont de Marsan, France; cKle = private collection of ANDREAS KLEEBERG, Berlin, Germany. Types - The interpretation of nearly all names mentioned in this work is based on the study of the relevant type specimens. More than 200 primary types were examined for the ingroups (including