Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Bálint, Zs.: Arcas Swainson, 1832 is revisited: review of some species-group names, identification of the sister group and a key for species (Lepidoptera, Lycaenidae: Eumaeini)

Remarks - In the CNB the species A. alleluia was erroneously lumped under A. nicolayi SALAZAR et COSTANTINO, 1995 (ROBBINS 2004), which I consider as one of the junior subjective synonyms of A. splendor (DRUCE, 1907) NlCOLAY, 1971 (see below). The species A. alleluia is the only Areas without dorsal fore wing androconia. The female either possesses a faint submedian pattern in the fore wing ventrum or she does not. In this character it is similar to that of A. delphia female, which is larger and has a very wide black margin in the dorsal fore wings. Consequently, I reinstate the nominal taxon alleluia to its original status. Areas arcadia BÁLINT, 2002, stat. n. Areas (tuneta) arcadia BÁLINT, 2002: 153, figs 17 (holotype dorsum), 18 (holotype ventrum), 19 (paratype female dorsum), 20 (paratype female ventrum). Areas tuneta (HEWITSON); ROBBINS 2004: 119 (synonymy in error). Material examined - BRAZIL: Santa Catharina (1 male, MNHN); Santa Catharina, Blumenau, 1910, Wernicke (1 male, 1 female, MNHN); no precise locality (1 male, 1 female, MNHN). Remarks - The material listed above displays the same characteristics that have been found to be diagnostic for the representatives of the superspecies A. tuneta living in the Atlantic region of South America: (1) wide black marginal border reaching dorsal fore wing androconia, and (2) golden basal scaling along ventral hind wing medial black line. Both semispecies of superspecies Rekoa zebina (HEWITSON, 1869) ROBBINS, 1991 were converted into species rank in the CNB without any comment (ROBBINS 2004: 123). Similarly, there is no explanation, why A. arcadia has been lumped under A. tuneta and why A. delphia NlCOLAY, 1971 is regarded as distinct species in the CNB. The phenomenon that allopatric species of Amazonian and Atlantic distribution exist is well known in butterflies (HALL & HARVEY 2002£>). Therefore, I consider the taxon arcadia as an Areas species in accordance with the methodology applied in the CNB and on the basis of the mentioned phylogeographic evidence. Areas delphia NlCOLAY, 1971 Areas delphia NlCOLAY, 1971: 101, holotype male: COSTA RICA, Guapiles, 850 Ft; Figs 21; BÁLINT 2002: 153, figs 21 (holotype dorsum), 22 (holotype ventrum), 23 (holotype labels). Areas katia SALAZAR et JOHNSON in JOHNSON & SALAZAR 2002: 148, figs 1 (holotype dorsum), 2 (holotype ventrum) 3 (genitalia); ROBBINS 2004: 119 (as A. delphia new synonym with no support); syn. n.