Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 98. (Budapest 2006)

Gattolliat, J.-L. ; Sartori, M.: The mayflies of Ghana (Insecta: Ephemeroptera)

the broader type. All specimens listed above also fit the broader morph concept, al­though the poor preservation of part of this material makes the estimation of the ra­tio of the turbinate eyes difficult. The generic attribution of the species to Pro­cloeon BENGTSSON, 1915 or to Cloeon is unclear; the species was originally de­scribed in the genus Procloeon (KlMMINS 1955) then moved to Cloeon (DEMOU­LIN 1970) and finally put back in Procloeon (GILLIES 1997). The situation will remain unclear until a complete revision of the genera Cloeon and Procloeon is un­dertaken. Distribution - Procloeon cylindroculum is widely distributed in Africa from Malawi (Nyasaland) in the south, to Tanzania in the east and Senegal in the west (GILLIES 1980b, KlMMINS 1955). In West Africa, it is reported from Gambia, Senegal and Ivory Coast (GILLIES 1980b). These first citations from Ghana are consequently included in the previously known distribution. Pseudocloeon cf. intone GILLIES, 1994 Baetis mtonis GILLIES, 1994 (orig.) Labiobaetis mtonis: LUGO-ORTIZ & MCCAFFERTY (1997) (comb.) Pseudocloeon intone: LUGO-ORTIZ, MCCAFFERTY & WALTZ (1999) (comb.) Material examined - GHANA: Ashanti Region, Kumasi, Volta, 1°36'W, 6°43'N, 293 m, 6 0*0*, 03.06.1967 (G224); same locality, 6 SS, 02.07.1967 (G230). All collected by S. ENDRŐDY­YOUNGA. Further material - IVORY COAST: Cavally, Road Tiboto-Grabo, 3 Sa, 11.02.1988, (A0263), J.-M. ELOUARD (ORSTOM) coll. and leg. Diagnostic features - Labiobaetis NOVIKOVA et KLUGE, 1987 was raised to the generic rank and included most species of the Baetis propinquus (WALSH, 1863), B. atrebatinus EATON, 1870 and B. molawinensis MÜLLER-LIEBENAU, 1982 species group by MCCAFFERTY & WALTZ (1995). Several African species previously assigned to Baetis were transferred to this genus (LUGO-ORTIZ & MCCAFFERTY 1997). Despite the dubious status of the genus Pseudocloeon KLAPÁLEK, 1905, all the species previously included in Labiobaetis and some spe­cies placed in Baetis LEACH, 1815 were subsequently assigned to Pseudocloeon (LUGO-ORTIZ et al. 1999). Because of the controversial status of the genus Pseu­docloeon and the lack of clearly shared diagnostic characters, GATTOLLIAT (2001) refuted the placement of the Malagasy species of Labiobaetis in Pseudocloeon. Re­cent molecular analysis show that the concept of Labiobaetis/Pseudocloeon is polyphyletic and that several lineages occur in the Afrotropical Realm (MONAGHAN et al. 2005). The examined material possesses the following characters: hindwings present with two longitudinal veins and without costal spur; sclerotized plate be-

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