Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 97. (Budapest 2005)

Bálint, Zs.: A review of the Neotropical hairstreak genus Annamaria with notes on further genera (Lepidoptera: Lycaenidae)

tinct in the examined species. The tegumen is large and heavily sclerotized, the saccus is long. The female genitalia is a membranous, less sclerotized organ. The ductus is relatively strong and wide, ventral membranous lateral plates surrounding the posterior antrum. The cervix is also membranous, but slightly sclerotized and joins the ductus to the bursa laterally. The bursal signa are wide, membranous and slightly pointed. Unique combination of diagnostic characters - ( 1 ) Very short male discal cell with scent pad in the discalis, (2) distinct scent patch in all intercellular space beyond the discal cell is present, (3) hind wing with three tailed vein termini plus an extended (lobated) anal margin in both sexes (restricted to the draudti lineage). This combination of characters has not been found in any other eumaeine lycaenids and seems to be unique. Character (1) is not unique in eumaeine lycaenids as male Micandra platyptera (FELDER et FELDER, 1865) possesses this trait or certain Panthiades HÜBNER, 1819 (type species: Papilio pelion CRAMER, 1775) species. However, there is no further trait which could support the tight relationship with any of the mentioned taxa, plus the discoidal scent pad of these genera are very distinct. Character (2) seems to be unique in the tribe; as some other eumaeine hairstreaks have also post discal scent patches, like Atlides HÜBNER, 1819 (type species: Papilio halesus CRAMER, 1777), Panthiades and Theritas HÜBNER, 1818 (type species: Theritas mavors HÜBNER, 1818), but the patches in these taxa are generally fused together and the scales are often qualitatively distinct in their morphology (cf. for Atlides figs 7-11 in BÁLINT et al. 2003). Character (3) seems to be also unique excluding the anal lobe, as for example Paiwarria KAYE, 1904 (type species: Papilio venulius CRAMER, 1779) is also "three tailed" but its hind wing anal angle is lobated like in Columbia and rhaptissima group, or in Ipocia BRÉVIGNON, 2002 (type species: Thecla sponsa HEWITSON, 1867). Therefore an analysis of wide array of related taxa (see Discussion) is necessary for identifying whether the character homoplasious or not and ancestral or derived in the case of the draudti group, plus in the whole genus. Systematic placement - Annamaria belongs to the tribe Eumaeini as characterized by ELIOT (1973: 439-440). It has ten fore wing veins, hairy eyes, a stubby tipped male fore leg tarsus and "greyhound-shaped" male genitalia lacking a juxta. The head frontoclypeus is without modified hair-like scales; ordinary scales directed downwards identify probably a broader monophyly which also includes Areas SWAINSON, 1832 (type species: Papilio imperialis CRAMER, 1775) and other conspicuous and well known eumaeine lycaenids. However, some of these taxa possess an extraordinary trait, namely a more or less developed ventral hind wing androconial pouch (ELIOT 1973: 402). Accordingly, I am of the opinion that all the taxa which possess this trait like Areas SWAINSON, Brangas HÜBNER, 1819