Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 86. (Budapest 1994)

Argaman, Q.: Generic synopsis of Myzinidae (Hymenoptera: Scolioidea)

ANNALES HISTORICO-NATURALES MUSEI NATIONALIS HUNGARICI Tomus 86. Budapest, 1994 p. 85-104 Generic synopsis of Myzinidae (Hymenoptera: Scolioidea) by Q. ARGAMAN, Bet Dagan ARGAMAN, Q. (1994): Generic synopsis of Myzinidae (Hymenoptera: Scolioidea). - Annls hist.-nat. Mus. natn. hung. 86: 85-104. Abstract - The classification of Myzinidae is reviewed and a key to genera is presented. Three new subfamilies (Iswarinae, Meriinae, Mesinae) and 13 new tribes: (Hylomesini, IUoswiini, Iswarini, Komarowiini, Macromeriini, Meriini, Mesini, Myzinellini, Parameriini, Poecilotiphiini, Warayoini, Weerpagini, Yooloini) are described, as are 23 new genera (Arapatka, Cocovasna, Ekepirka, Fikoplesa, Fongiba, Fukpokta, Gonordula, Hahiya, llloswia, Ivazuga, Jurja, Keyovaska, Nurmiya, Nyuka, Taywala, Tilkuya, Tokoparta, Upaterka, Warayoa, Weerpaga, Xilunka, Yooloa, Zezelda) and 3 new species (Jurja limpida, Weerpaga udomanca, Yooloa vircola). A replacement name, Warayoa citreosigna, is proposed for Braunsomeria quadraticeps TURNER, 1912 male only. Neotypes are selected and redescribed for Myzine arcuata and M. spinosa. With 89 figures. INTRODUCTION The family Myzinidae, as conceived here, is differentiated from Tiphiidae, with which it was united once by PATE (1947), by the following apomorphous character states: 1. Antennae apparently 12-segmented in male and 11-segmented in female, since the pedicel is invaginated into the apex of scape. 2. Antennái toruli covered by (or entirely removed from plane of front) the strongly developed antenna! tubercles. 3. Propodeal spiracles situated close to base of the unbordered propodeal disc, perpendicular to metanoto-propodeal suture. 4. Prepectal carina of mesopleuron not fully developed in either sex, although it is often traceable through a delicate ridge. 5. Scape of male twice longer ventrally than dorsally. 6. Marginal cell of female forewing, when developed, removed from the costa. 7. Female hind tibia without sensorial pit, instead the longer spur modified into a calcar. For the corresponding character states in Tiphiidae see ARGAMAN & ÖZBEK (1992). KROMBEIN (1937) noticed the subsequent separation of Myzinidae from Brachycistidinae (Tiphiidae). Some representatives of Thynnidae, however, share the same reduced number of antennái segments as it occurs in Myzinidae. Separation of Myzinidae from Thynnidae will be discussed elsewhere. The supraspecific nomenclature of myzinids has its own intrinsic problems. Generic-group names of RAFINESQUE were considered lapses or emendations for a long time. KROMBEIN (1937), however, designated Tiphia tripunctata ROSSI, 1790 as type of Meriana RAFINESQUE, 1815, thus validated it. KROMBEIN'S intention was to sink Meriana, making it isogenotypic and synonymous with Meria ILLIGER, 1807. At that time, latreillei FABRICIUS, 1804 was considered a junior synonym of tripunctata, and the latter, due to the synonymy, a type of the genus Meria. As a result of the present study, I not consider latreillei and tripunctata to be synonymous or congeneric. Hence the type of Meria is not tripunctata but latreillei by monotypy, Meriana becomes a valid genus with tripunctata as type. Moreover, in virtue of KROMBEIN'S (1937) validation, other names proposed by RAFINESQUE: Tiphiana and Myzina (latter with its first included species, hispanica, by SPINOLA in 1843, as type), are regarded to be valid as well. The name Meira of CAMERON (1902), in combination with the species quadrimaculata CAMERON, 1902, but without an explicit indication that he intended to propose a new genericgroup name, was surely an ordinary lapse frequently found in the notes of that author, which must be rejected.