Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 86. (Budapest 1994)

Bálint, Zs. ; Johnson, K.: Polyommatine lycaenids of the oreal biome in the Neotropics, part II: The Itylos section (Lepidoptera: Lycaenidae, Polyommatinae)

APPENDIX - ADDITIONAL NOTES ADDED IN PROOF Dr. GERARDO LAMAS, Director of the Museo Nációnál de História Natural de la Universidad Nacional Mayor de San Carlos (Lima, Peru) [MUSM], has collected extensively throughout Peru and discovered several lycaenid butterflies new to science. Some of these were reported in the literature (e.g. LAMAS & PEREZ 1983, Figs 37-43) but not formally described. After publication of BÁLINT'S (1993a) catalogue of Neotropical Polyommatinae, Dr. LAMAS provided numerous comments and also mentioned that the MUSM butterfly collection included some additional undescribed entities from the genera Madeleinea BÁLINT, 1993 and Itylos DRAUDT, [1921] (BÁLINT 1993b). Formal descriptions of these taxa are currently being prepared by the senior author and Dr. LAMAS. Since the aim of the present paper was to revise the Neotropical Itylos clade, it is important to briefly summarize results which will be published subsequently by BÁLINT & LAMAS and add comments pertinent to this revision. Taxonomic description of a new Itylos species The following entity will be formally described by BÁLINT & LAMAS and, in context with the taxa of the present revision, should be looked for. Diagnosis (Fig. 67) - Superficially most similar to /. pnin BÁLINT, 1993 but with VHW postmedian band more prominent. Male genitalia (Fig. 68) fitting the clade including /. pnin and /. titicaca (WEYMER, 1890) but with male genital uncus and gnathos more elongate and slender, aedeagus shorter and with large bulbous vesica. Female: Unknown. Current data - The species is currently knwon from a single male labelled "Peru, PA, km 300, Carrt. Lima-Huánuco, 4300 m, 31.V.'78., G. LAMAS", genitalia dissection 396, gen. prep. No. BÁLINT, in glycerin vial, deposited in MUSM. Dr. LAMAS notes that the locality as "4300 m, Pasco, 300 km Carretera, Lima-Huánuco, Peru". Female genitalia of Itylos pnin The original description of /. pnin was based on a single male specimen misidentified by former NHM curators as female /. titicaca (BÁLINT 1993a). Subsequently Dr. LAMAS located a female specimen (Fig. 69) at the MUSM which he and the senior author have identified as the female of /. pnin. The association is based on (1) similar wing shape; (2) identical coloration of antennae; (3) identical VHW pattern and (4) collecting locality in same region as type locality of /. pnin. The specimen is labelled as follows: "Pern, Huánuco, Pachas, 3450 m, 23.VIII.1965, P. Hocking"; genitalia dissection is gen. prep. No. 399, BÁLINT, in glycerin vial, deposited in MUSM. Female genitalia (Fig. 70) - In dorsal view, the henia are heavily sclerotized and the fibula bilobed; the apical lobes long and rounded, the dorsal lobes wide and pointed and the central element tubular. In ventral view, apical fibular process is elnogate (quite equal to length of anterior lamella) and slightly curved; the ductus bursae is short (about the same length as the henia) and the anterior apohysis of the eighth tergit short and stout. Comparative Remarks Phylogenetically, the new Peruvian Itylos species fits well into the character transformation series we have previously elaborated (see "Review of Ityloides BALLETTO, 1993", above). The new Peruvian Itylos appears to be the sister species of /. pnin and these sister couplet of /. titicaca. The unique female genitalic structures of /. pnin indicated corroborate our proposal to remove Itylos from ELIOT'S Polyommatus-sectlon and create an additional section. Female genitalia typifying the Polyommatus section show a relatively long ductus bursae (three or more times longer that that of henia) which is evertable. The ductus bursae of Itylos is not evertable