Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 86. (Budapest 1994)

Bálint, Zs. ; Johnson, K.: Polyommatine lycaenids of the oreal biome in the Neotropics, part II: The Itylos section (Lepidoptera: Lycaenidae, Polyommatinae)

oi Paralycaeides shade, Madeleinea lolita and M. mashenka in BÁLINT 1993a). This species is described almost a hundred years after the date of its capture and suggests that numerous "hidden" entities may still exist in institutes or museums where historical South American polyommatine collections were preserved. The description of /. fumosus and /. luzhin is based on the same specimens (BÁLINT 1993b: 1-2). Type Material Examined - Holotype, male, labelled as follows: "Paramo, Coreuges, Peru, 36004000 m, 30.XI.99., dry seas. (Simons); Rothschild Bequest, B.M. 1939-1.; Holotypus, Itylos luzhin, det. Zs. Bálint, VIII. 1992., London". - Paratypes, 2 males, with the same data. Deposited in BMNH (drawer 29A-928). Slides: Brit. Mus. Nos. 17569 (Holotype); 17570, 19170 (Paratypes). Etymology - "Fumosus"; an adjective indicating the fumous DW and HW ground of the wings. "Luzhin"; named for the chess monomanic in NABOKOV'S novel "The Defense". Review of Ityloides BALLETTO, 1993 Short Taxonomic History - BALLETTO (1993: 232, 234) describing Ityloides fumosus used the supposed new entity as type species of a new monobasic genus Ityloides. Some misfortune resulted here because of arbitrary dates of publication and the fact that little communication had occurred concerning the preparation of research on these entities. The publication of BÁLINT was tied to names needed for a Chilean publication (L. PENA in litt, to JOHNSON). BALLETTO was unaware of BÁLINT'S treatments of the name Itylos, titicaca, speciosa and of the existence of /. pnin. Ityloides, however, is a published name and requires evaluation in regard to a revisionary view of Itylos. As noted above, we synonymize monobasic Ityloides with Itylos for the factual and methodological reasons enumerated below. Diagnosis of Ityloides- Body: eyes naked, palpy hairy, foreleg tibial spine absent, abdomen short. Wing venation: FW with veins Sc and Rl free (Fig. 66). Wing shape: FW with sligtly convex, apex pointed, inner margin short and convex, tornus rounded, anal margin very short comparing with costa. Pattern: DW ground deep smoky indigo blue with wide suffused black margin. FW with discoidal spot. Fringes brownish, fumosed, type 7. FW costa fringed at apex. VFW ground greyish brown with suffused polyommatine resembling markings. VHW pattern ityloid: ground colour greyish brown with darker discoidal spot, postbasal and postmedial row; costal area with submedian dark brown spot, black median and postmedian pattern; submarginal pattern indistinct, suffused. Androconial scales: absent. Male genitalia: whole dorsum (uncus + gnathos + tegumen) remarkably similar in type to polyommatine lycaenids, characterized by well developed uncus, smaller gnathos and stronger tegumen; uncus digitate with about the same lentgh as juxta, straight with bulbous, rounded apex, apical lobe dentated; ganthos shorter than uncus (about 5/6 length of that of uncus) and pointed with strongly shouldered base; tegumen and vinculum with about the same length, cca. equal with the length of aedeagus' subzonal part, appendix angularis absent; juxta saddled and slightly curved invardly, somewhat shorter than the subzonal part of the aedeagus; with short but well developed arms and pointed apex; valva wide with strongly angulated but rounded costa, rostellum thick but strongly dentated, anal margin sligthly convex, strongly sclerotized with pointed apex; aedeagus straight, long, suprazonal length about one third as that of subzonal; suprazonal part thin with extanded sclerotized lateral edges (CHAPMAN'S process), pointed at apex, about two third longer than vesica; vesica very simple and weak; subzonal part cylindric; alulae very small, sagum absent. Female: Unknown. Synopsis of the Species - Ityloides was monobasic, including only Ityloides fumosus BALLETTO, 1993. Evaluation of Status - Characters of Ityloides: Ityloides was characterized as showing (1) strong, bulbous uncus apex with dentated lobe, (2) thin, long and dentated costal process of valve, (3) penis with screlotized and straight suprazonal edges, (4) very week alula. These genitalic structural characters are unique to /. fumosus. However, there is no evidence at this time concerning which of the unique chararacters of /. fumosus are apomorphic. /. pnin has also bulbous uncus but without dentation; the sclerotized edges of suprazonal part of the penis are also present in /. titicaca although the structure is curved and longer compared to the subzonal portion. The valval rostellum on /. fumosus cannot be compared with any polyommatine occuring in South America. However, the alulae of /. titicaca and /. pnin are conside-