Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 86. (Budapest 1994)

Bálint, Zs. ; Johnson, K.: Polyommatine lycaenids of the oreal biome in the Neotropics, part II: The Itylos section (Lepidoptera: Lycaenidae, Polyommatinae)

treated as available by numerous authors and listed again in Itylos (sensu NABOKOV!) (cf. DESCIMON (1986, Tab. 20-3) or as Parachilades titicaca speciosa in BRIDGES (1988: 1.326, 11.81). Because of this, BÁLINT (1993a) tentatively listed the specific status of speciosa again, and placed it and titicaca in Itylos until the type of titicaca will be chequed. While the mentioned paper of the senior author was in press, one syntype specimen of titicaca (with a round "syntype" label added by Dr. LAMAS) was sent to him for study. Accordingly, this specimen has been designated lectotype of Lycaena titicaca below (Figs 11, 12, 14). Facies of this type strengthen the opinion of NABOKOV: the taxa titicaca and speciosa appear identical and we here designate Cupido speciosa STAUDINGER as a junior synonym of Lycaena titicaca WEYMER, syn. n. Sex Recognition: Sexes of/, titicaca can be distinguished by FW ground colour and shape (details above in diagnosis of the species). Intraspecific Variation: Some female specimens have strong bronze shade DW ground. The VFW ground is ash grey in some cases (Fig. 6). The postdiscal spots in the upper postmedian area are differently developed on several specimens: sometimes rather distinctly marked (Fig. 4), sometimes totally missing (Figs 6, 8). VHW is rather variable, the two most often phenomena: (1) the area between costal margin and vein Ml is paler (Fig. 2); (2) the ground of the median area is dark creating a wide brown band with the postbasal and postmedian markings (Figs 4, 6). In genitalia, only males show differences in development and shape of costal valval rostellum and anal process (Figs 50-52). Female genitalic structures are rather uniform according to the studied material. Subspecies-like demes: Two stocks of material indicate the possibility of subspecific division, but the available material is too small to render a conclusive diagnostic treatment. We briefly characterize these entities below: 1) deme of Ancash-Cordillera Bianca (Figs 1-2, 17-20, 35, 41, 58). Material Available: 6 males (details in Material Examined). Additional characters: slightly different wingshape and larger size; DW ground gleaming purple blue; DHW ground fuliginous grey. 2) deme of Tucumán-Antofagasta (Figs 15, 16, 42, 60). Material Available: 13 males 3 females (details in Material Examined). Additional character: valval anal tip straight, spinous (Figs 42, 60). There are no elinal changes toward these phenomena and the mentioned characters seem to be unique compared to the overall range of titicaca. Both entities also show distinctive geographic positions as disjunct populations peripheral to the centrally documented range of titicaca (Fig. 33). It is probable that when more material becomes available for study distinct subspecific status of these entities might be advised. Interestingly, the exsistence of the phenomenon Ancash was suggested before by NABOKOV (see BÁLINT 1993a: 13). Phylogenctic Affinities and Placement: The taxon seems to be the sister species of /. pnin. (for additional details see below under Discussion and Appendix). General Comments-/, titicaca, beside Pseudolucia chilensis (BLANCHARD, 1852), P. lyrnessa (HEWITSON, 1874) (= zembla BÁLINT et JOHNSON, 1993a: 6-7) and Madeleinea koa (DRUCE, 1876), is the most well represented South American polyommatine taxon in the European collections (cf. BÁLINT 1993a). As the types of titicaca and speciosa were never studied and compared hitherto, up to the very recently the taxon was often determined as Itylos speciosa in European museum collections (London, Paris, Munich, Vienna) following the misinterpretation of DRAUDT ([1921]: 822). Type Material Examined - Lectotype of Lycaena titicaca, male (Figs 12, 14), labelled as follows: "Zool. Mus. Berlin; Syntype; Lectotypus, Lycaena titicaca, des: Zs. Bálint, 1993.III.3., Budapest". Deposited MNHU; designated here. Lectotype of Cupido speciosa designated by BÁLINT (1993a: Fig. 6), male: "Malaga, Bolivia, 4-5000, M. Garlepp '91.; Speciosa Stgr.; Origin.; Zool. Mus. Berlin; Syntype; Lectotypus, Cupido speciosa, des: Zs. Bálint, 1992.XII.2., Budapest". Deposited in MNHU. According to the description of STAUDINGER (1

894: 77-78) the type series of Cupido speciosa, contains at least