Matskási István (szerk.): A Magyar Természettudományi Múzeum évkönyve 83. (Budapest 1991)

Kvaček, Z. ; Hably, L.: Notes on the Egerian stratotype flora at Eger (Wind Brickyard), Hungary, Upper Oligocene

TETRACENTRACEAE Tetracentron agriense (ANDREÁNSZKY, 1962) comb. n. (Plate IV: 1-4) B a s i o n y m : Acer agriense ANDREÁNSZKY 1962, 230. pl. 3. f. 3. textf. 7. Diagnosis emend.: Leaves orbicular ovate, petiolate, 8-12 cm long and 3.5-6.0 cm wide (some fragnentary specimens of obviously larger size). Base slightly cordate to rounded, apex bluntly acuminate. Petiole fragment 33 mm long. Venation palmate, semicraspedodromous. Basal primaries 5-7, secondaries steep (the angle of divergence 20°-30°), looping along the margin, side veins forming another row of small loops, from which small veinlets enter the teeth. Tertiary veins dense, mostly percurrent, spidernet-like arranged. Margin simply coarsely crenulate, teeth glandular, blunt, sinuses rounded to widely cuneate. Hoi o type: MM 76.11.2 (ANDREÁNSZKY 1962, textf. 7) - EWU (with its counterpart). - Paratype s : MM 61.811.1 (W1559) (ANDREÁNSZKY 1962, pl. 3, f. 3) 61.812.1 (W 1325/a), 61.821.1 (W 13261a), 62.114.1, 62.189.1,62.312.1 - all EWU. A superficial resemblance of these leaves, which are rarely met in EWU, to Mafvales is misleading. The glandular teeth exclude such a possibility. The picture given by ANDREÁNSZKY (1962, textf. 7) does not reflect true course of side veins, which are looping along the margin. In the marginal venation and teeth we find connecting features to Trochodendrales. The venation patterns best match with the extant Tetracentron sinense OLIV., which differs in finer teeth and attachment of the petiole. In the fossil species the petiole is attached from the underside while in the recent one it lies in the same plane with the leaf lamina. Of the so far described fossil Tetracentron remains, the leaves of Tetracentron hungaricum ANDREÁNSZKY (1959b) from the Sarmatian tuffs at Balaton (Hungary) differ in the nearly entire margin, but we were unable to study the type specimen, which was missing from MM. The Cretaceous Tetracentron potomacense (WARD) ILJINSKAJA is very deeply cordate and finely crenulate, T. walirameevianum IUINSKAJA differs in its very small size, the arrangement of basal primaries (see ILJINSKAJA 1972). We found the same kind of attachment of petiole in the newly described Tetracentron beringianum CELEBAEVA et SANCER (in press) from the early Palaeogene of the Far East as well as in some species of Trochodendroides (T. vassUenkoi ILJINSKAJA et ROMANOVA). Tetracentron sinense Ouv., a small deciduous tree of eastern and south-eastern China grows in mesophytic subtropical forests, and we can expect similar autecology of T. agriense. PLATANACEAE Platanus neptuni (ETTINGSHAUSEN, 1866) BÖZEK, HOLY & KVACEK, 1967 (Plate V: 4) Typical oblong, bluntly simple toothed leaves with semicraspedodromous venation are more frequent only in EWL (about 20 specimens at MM) but quite rarely in EWM (MM 62.658.1, 64.39.1, 65.704.1). As already stated by HABLY (1985) they were described as Cunonia oligocenica ANDREÁNSZKY et NovÁK (1957). ANDREÁNSZKY (1966) identified this type of leaves not only with C. oligocenica (I. c. textfig. 77, MM 61.816.1) but also as Pterocarya denticulata (O. WEBER) HEER (MM 64.73.1, 64.77.1) and Quercus legányi ANDREÁNSZKY et KovÁTS (MM 64.39.1). P. neptuni is a common plant of the Hungarian Egerian and Kiscellian. Its occurrence was proved cuticular study (HABLY 1980). ULMACEAE Ulmus fischeri HEER, 1856 (Plate V: 2) Large oval, strongly asymmetrical, partly deeply cordate leaves attaining a length of more than 10 cm, usually coarsly double serrate, in smaller specimens with simple dentation, represent the group of elm leaves, called by ANDREÁNSZKY Ulmus sp. type I. (ANDREÁNSZKY 1966, textfig. 60, 61). This species represents a