Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 71. (Budapest 1979)

Balogh, J. ; Mahunka, S.: New taxa in the system of the Oribatida (Acari)

(Euarthronota) to include all other arthronotic Oribatids within the new supercohors Arthronota. The 10 families of the new cohors belong in 8 superfamilies. Of the 8 superfamilies 7 have only one family each and of these families only the Brachychthoniidae BALOGH, 1943, has a number of genera and species. The third subfamily Phyllochthonoidea TRAVÉ, 1967, includes 3 families, all of which are monotypical. Supercohors 3: Parhyposomata supercohors nov. contains highly similar Oribatids of a parhypochthonoid organization. The very few species of this group belong in 2 families representing only one superfamily. Supercohors 4: Mixonomata GRANDJEAN, 1969, seems a very heterogeneous taxon: it should perhaps be further subdivided in the future. For the time being we consider as necessary the separation of merely 2 cohors. All holonotic and dichoidic taxa of the Mixonomata are relegated to the first cohors, Dichosomata cohors nov., whereas the holonotic and ptychoid forms belong in the other one, Euptyctima GRANDJEAN, 1967. Both terms, Dichosomata and Holosomata, as well as their definitions were given in 1969 by GRANDJEAN, although without designation of category status. Following his concept and developing it further we deem it reasonable to erect the two supercohors mentioned above as Dichosomata and Holosomata. In the new cohors Dichosomata belong 5 superfamilies, each with merely 1 family in the Palaearctic Region. The other cohors, Euptyctima, includes only 2 superfamilies with 4 families. Supercohors 5: Holosomata supercohors nov. It includes all Oribatids with a sclerotized sejugal tegument. The new supercohors contains 3 superfamilies with 6 families. The short diagnoses of the subordos, supercohors and cohors in the Oribatid system as here elaborated are as follows : Archoribatida subordo n. One or many characters are of the most primitive type. Hysterosoma sometimes archeonotic without notogaster; shield separated, eventually partly or wholly absent, or separated by wide intermediate spaces. Under such circumstances body weakly pigmented, whitish, sometimes with large, movable, plumose black hairs. Legs hexadactylous, with 2 femora. Species belonging in other groups with an arthronotic notogaster of 1-3 tranverse suturae, with 2-4 tergites, or holotonic, i.e. undivided. Anogenital region mostly macropolic, that of in most ancestral forms heavily divided; genital, aggenital, anal, paraanal, adanal, praeanal and postanal plates may be present, but during development eventually variously fused. Ptychoidy, dichoidy and holoidy may be equally present. Brachypilic anogenital region occurring very rarely and together with pseudogastry or schizogastry ; hologastry known only in one species. The position of hairs on the genital plate reveals the primitive phylogenetical status of the new subordo. Euoribatida subordo n. Without any primitive characters. Body always pigmented. Dorsal lamellae as well as ventral ones fused into a single huge plate (prodorsal, notogastral, epimeral and anogenital lamellae). Neither ptychoidy nor dichoidy, only holoidy occurring. Anogenital region brachypilic. Legs pentadactylous.