Kaszab Zoltán (szerk.): A Magyar Természettudományi Múzeum évkönyve 65. (Budapest 1973)
Ujhelyi, J.: Data to the systematics of the sections Bulbosae and Caespitosae of the genus Koeleria, XI.
schinzii (DOM.) UJH. — are bulbous irrespectively of origin, while all others are cespitose. I have brought forth examples in several series. Similarly, the compact or loose nature of the panicles displays the correlation between the species and its habitat conditions (ÚJHELYI, 1961—1972). Deducing the "hybrids" from a species each of the bulbose or cespitose sections, DOMIN regarded the phenomenon as an egregiously interesting botanical phenomenon. Of the first section, the "hybrid" unquestionably participates in Koeleria hirsuta (LAM. & DC) GAUD., as to the colour, shape and repent state of the leaves, the coloured spikelets (they are more pallid than those of the original ones living in the subalpine region), the pubescence of the upper culm and glumae, and the short, aristate paleae. The main feature of the Sectio Bulbosae, the incrassation of the base, had (according to DOMIN) been lost. He looked for the other parent in the Sectio Cristatae, thinking of "Koeleria pyramidata ' or the "highly variable Koeleria gracilis". One of these latter parents had, in his opinion, influenced growth conditions (DOMIN alludes here to the thin culm basis). These were presumably responsible for the pubescence of the blade, the loss of indûment of the spikelets, and the shortening of the aristae. I have grown so many of DOMIN'S plants relegated to the Sectio BULBOSAE in my uniform garden during the last two decades that I am in a position to answer all of his theoretical inferences. Unfortunately, I had until now no opportunity to grow Koeleria hirsuta (LAM. & DC) GAUD., or Koeleria schinzii (DOM.) UJH., but I was able to observe every member of similar ecological requirements and alliance of the Series Setaceae UJH., the Series Gaudatae ÜJH., as well as Koeleria brevifolia REUT., in my experimental garden. Despite the fact that all of them have retained their genetical features —-thus the vetust vaginae of the species assigned to the Series Setaceae UJH. remained reticulate, not has the anatomical structure of the leaf epiderm essentially changed —the incrassation of the base, appearing pregnantly in the original habitat, failed to occur and the vetust vaginae elongated to an extent characteristic of the species. On the dry, insolated alpine meadows, on the cliffs or in their crevices or on the stone-rubbles, at altitudes of 2000-3000 m and by the added effect of the ultraviolet radiation, the vaginae are naturally hardly 1—2 cm long, but in the alpinetum of the 1200 m high Pont du Nant the environmental conditions were naturally different. The pubescence of the innovational leaves and the glabrous spikelets, as well as the shortening of the aristae, are by far not proofs of a hybridous descent. HOFPE'S valuable collection was included in DEGEN'S herbarium presented to the Department. One of its sheets contains the following data: "224. Koeleria hirsuta GAUD. In rupibus alpium editissimarum Tyrolis australis. Julio." Besides a plani with 1 cm short vaginae, there is one with 3 cm, thus not shorter than those of Koeleria Wilczekiana DOM. Among the hirsute spikelets, there are also wholly glabrous ones. I found specimens with glabrescent spikelets also in other collections. Incidentally, the base of the holotype of Koeleria schinzii (DOM.) UJH., the descendant of Koeleria hirsuta GAUD., is not incrassate, and the vaginae attain a length of 3 cm. Beyond the macroscopic features, the anatomical investigation of the plant dispersed every doubt. True, most specimens of Koeleria hirsuta GAUD, are glabrescent, and appears glabrous even when observed through a lupe of small magnification, they are only juvenile blades. Owing to the strong innovation, the sensile leaves become oppressed, yellowish, hence when only a fragment is torn from the lawn, the herbarial
