L. Forró szerk.: Miscellanea Zoologica Hungarica 7. 1992 (Budapest, 1992)

Topál, Gy.; Csorba, G.: The subspecific division of Rhinolophus luctus Temminck, 1835, and the taxonomic status of R. beddomei Andersen, 1905 (Mammalia, Chiroptera)

p. 121), although they remarked "their treatment as races is provisional"..and ..."A detailed and painstaking analysis with a large quantity of material will be required before the races of R. luctus can be worked out satisfactorily" (Tate 1943 p. 5). It is somehow strange that Tate (1943), while retaining Andersen's original "groups", put R. pearsoni in the R. luctus group and thus - in a later sense - as a subspecies of the latter. Well after that, R. pearsoni was recognized as a clearly distinct species by Ellerman & Morrison-Scott (op. cit.). (Incidentally, the specimen of R. I. pemiger deposited in the Hungarian Natural History Museum, was found in the same cellar as a small colony of R. pearsoni at Tarn Dao in Vietnam by the present junior author.) Sinha (1973) gave details on the material in the Calcutta collection of the Zoological Survey of India. Lekagul & McNeely (1977) considered "probably two subspecies in Thailand: R. I. luctus in the south as far north as Tenasserim, and perhaps R. I. pemiger in the north". Payne et al. (1985) briefly discussed the ecology and habitat of R. I. foetidus and reported it from lowlands up to 1600 m in the mountains of Borneo. Liang & Dong (1984) and Chen et al. (1989) reported the species from further localities in Southern China. Ando et al. (1983) studied the karyotype of the species from Taiwan, Narayana Naidu & Gururaj (1984) in India, and Harada et al. (1985) the same topic in the specimens from Thailand. Materials and methods During the rather intensive collectings of bats by the senior author in India, this bat (R. beddomei) was seen but once in Southwestern India. The species and the South-Indian R. beddomei were studied briefly (by the senior author) in the Bombay Natural History Society's collection where there were available 6 specimens of R. I. beddomei, 6 specimens of R. I. pemiger and 1 specimen of R. I. lanosus in 1967. The female R. I. pemiger obtained by the junior author is the third known specimen from northern Vietnam, the first two were collected at the same locality and deposited in the Institute of Systematics and Evolution of Animals, Krakow, Poland (Cao Van Sung in litt.). Skulls of thirty specimens of Rfrinolophus luctus (s.l.) were used for the present study. List of the specimens with names of subspecies (R. I pemiger, R. I. morio, R. I. foetidus, R. I. beddomei), serial number for the present study, location of the specimen (Hungarian Natural History Museum, Budapest = HNHM, The Natural History Museum, London = BNHM, Bombay Natural History Society, Bombay = BNHS), register No., sex (male = m, female = f, undetermined = s?), and collecting locality as follows. R. 1. perniger: LHNHM 11111, f, Tarn Dao, Vietnam; 2:BNHM 78.2310., m,Chiangmai, Thailand; 3:BNHM 7.1.1.294., s?, "Calcutta", India(?); 4:BNHM 9.10.11.2., s?,Chiangmai, Thailand; 5:BNHM 21.1.6.4., m, Khonshong, Jaintia Hills, Meghalaya, India; 6:BNHM 21.1.6.5., m, Khonshong, Jaintia Hills, Meghalaya, India; 7:BNHM 79.11.21.142., s?, Masuri (Mussoorie), Uttar Pradesh, India; 8:BNHM 79.11.21.141., m, Masuri (Mussoorie), Uttar Pradesh, India; 9:BNHM 9.1.4.11., m, Darjeeling, West Bengal, India; 10:BNHM 91.10.7.55., s?, Sikkim; 1LBNHM 23.1.9.1., f, Chalma-Khel, Nepal; 12-.BNHM 21.1.6.2., Bankochori, S.Tenasserim, Burma; 13:BNHM 21.1.6.3., Kindat, Chin Hills, Burma; 14:BNHM 50.396., f, Nam Tamas Valley, Upper Burma; 15:BNHM 50.397, f, Taron Valley, Upper Burma; 16:BNHM 21.1.6.1, m, Sokteik, N.Shan State, Burma; 17:BNHS 3073, f, Khonshong, Jaintia Hills, Meghalaya, India; 18:BNHS 3072, m, Khonshong, Jaintia Hills, Meghalaya, India; ;9:BNHS 3071, f, Bouzini, Nepal. R. 1. morio: 20:BNHM 1.3.9.3, s?, Semangko Gap, Selangore, Malaysia; 2LBNHM 78.2309, f, Pak Thengchai, Sukerat, Thailand; 22:BNHM 70.1463, f, Korat Pn, Thailand. R. 1. foetidus: 23:BNHM 76.9.20.12, s?, N.W.Borneo; 24:BNHM 92.2.7.3, f, Mt.Dulit, Borneo; 25:BNHM 94.9.29.4, s?, Mt.Dulit, Borneo; 26:BNHM 98.11.3.9, s?, Lawas, Borneo; 27:BNHM 59.183, f, Lobang Badak, Serabang, Borneo. R. 1. beddomei: 28:BNHM 12.11.28.5, m, Sirsi, N.Kanara, Karnataka, India; 29:BNHM 11.3.16.1, f, Konkan, Maharashtra, India; 30:BNHS 3081, m, Karla Caves, Pune, Maharashtra, India.