Ábrahám Levente (szerk.): Válogatott tanulmányok VI. - Natura Somogyiensis 19. (Kaposvár, 2010)

HORVÁTH GY., HERCZEG R., TAMÁSI K. & SALI N.: Nestedness of small mammal assemblages and role of indicator species in isolated marshland habitats

294 NATURA SOMOGYIENSIS year we performed the IndVal analysis on the basis of data derived from 6 habitat patches of 4 macro-habitats. We recorded 13 small mammal species with the trapping of 6 habitat patches and 6 of them received significant value (Table 4). The yellow-necked mouse (A. flavicollis) got the highest IndVal value, however, despite of this fact its sig­nificant indicator value was expoundable only for that patch of Keleti-berek (KE_1), where human activities altered considerably the structure of vegetation in the previous years (Fig. 4). Furthermore, we received maximum indicator values for two other spe­cies (A. agrarius, M. minutus), but it was only the harvest mouse whose IndVal value was significant, which can be jointly construed for two habitat patches (H l, KE 2) on the second level of hierarchy. Therefore, the harvest mouse was a component of small mammal assemblages of several habitat patches as an asymmertic indicator species. Miller's water shrew (N. anomalus ) showed up as an asymmetric character species of one habitat patch (H_2) with significant maximum value, which indicated the higher level of water on this sedgy-reedy transient habitat mosaic (Fig 4). The european water vole (A. terrestris) with its 60% IndVal value can be considered as a symmetrical indica­tor species (> 55% IndVal) (Table 4). Its significant maximum character value character­ized the second habitat patch of Keleti-berek (KE_2) on the fifth level of the hierarchy, where the water level of the homogeneous sedgy area was higher in springtime (Fig. 4). Trapping throughout the year 2006 cannot prove the permanent presence of root vole subpopulation in the Halász-rét or rather its colonisation in the area of Keleti-berek. In this year the other endangered vole species, the field vole (M agrestis ) - that character­ises the Kis-Balaton marshy relict area - appeared in the second habitat patch of Keleti­berek (KE_2). There had been no human disturbance through the preceding few years, so the field vole got significant maximum IndVal value for the typical homogeneous sedgy habitat (Fig. 4). Table 4: Table of the indicator values of the species based on the hierarchy level of the habi­tat patches in 2006, where the first value is the number of caught individuals by single habi­tat patch and the second value is the number of those trap points where the species has occurred in the given patch. ,. . „ Keleti-berek Halász-rét Balatoni-berek Simon-sziget Sampling plot fadVal (%) Species KE_1 KE_2 HI H_2 B_3 S_1 S. araneus 50.77 6/5 41/13 18/13 0/0 4/2 0/0 S. minutus 11.76 0/0 2/2 0/0 0/0 0/0 0/0 C. suaveolens 7.41 0/0 0/0 2/2 0/0 0/0 0/0 N. anomalus 37.37** 0/0 18/9 5/4 2/2 0/0 0/0 N. foediens 7.41 0/0 0/0 2/2 0/0 0/0 0/0 M. agrestis 35.41** 0/0 12/10 11/7 0/0 1/1 0/0 A. terrestris 58.56* 0/0 60/11 10/4 0/0 0/0 0/0 R. rattus 7.41 0/0 0/0 3/2 0/0 0/0 0/0 A. agrarius 70.77* 3/3 262/16 89/21 2/1 14/3 14/2 A.flavicollis 84.87* 1/1 3/3 0/0 0/0 0/0 3/2 A. sylvaticus 9.09 0/0 2/2 2/2 0/0 0/0 0/0 M. spicilegus 29.41** 0/0 5/5 0/0 0/0 0/0 0/0 M. minutus 74.55* 0/0 46/16 85/21 8/3 0/0 0/0 Total 10/9 451/87 227/78 12/6 19/6 17/4 **:P <0.01; *:P< 0.05

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