Folia Historico-Naturalia Musei Matraensis - A Mátra Múzeum Természetrajzi Közleményei 17. (1992)

Bužek, Č.–László, J.: Contribution to the Upper Pannonian flora from Visonta, northern Hungary

CERATOPHYLLÄCEAE Ceratophyllum dubium (LUDWIG) KIRCHHEIMER (PI. II, Figs. 5-11): Description: Fruits (with or without exocarp) and their dehiscent halves, usually flattened, oval to long-oval in outline, 2.730 to 4.230 mm long and 1.455 to 2.270 mm wide, in some cases somewhat obliquely truncated at the base and with short slopping prickle bent dorsally, in apical or slightly subapical position; if prickles missing, usually not broken off, one specimen with stump of stronger apical prickle and with slight indication of basal (lateral) prickles; surface sculpture of exocarp smooth with or without dispersed minute warty-papillae, surface of endocarp smooth to finely granulated or paplllated, as well; inner side of some fruit halves with adhered remnant of the seed testa showing a conspicuous "mark" of the vascular bundles which is as characteristic for Ceratophyllum as the fruit vascularisation well seen in the plane of dehiscence. Remarks: Fruit remains well agree in all respects with those of the species Ceratophyllum dubium known from the Miocene of Europe, which is matched with the living species С submersum I. According to characters of the material studied fossil species seems to be closely related to C. submersum subsp. submersum var. submersum. Similar material but without apical prickles (if not broken off) was described from the Pannonian and Pontian of southern Moravia as C. dubium (CTYROKf and KNOBLOCH 1976) and C. protanaiticum (NIKITIN) 00R0FEEV (KNOBLOCH 1901a). The relation between this material and tuberculateri or prickled specimens described as C. cf. tanalticum SAPEGIN from the same stratigraphical level and region is not clear. But It must be noted here, that the fruit ornamentation in Ceratophyllum is very complex and variable (HILMOT-OEAR 1905). The living species C. tanalticum was recently banished to the synonymy of С submersum ssp. muricatum (Cham.) WILM0T-0EAR var. echinatum (A. GRAY) WILMOT-DEAR. Nevertheless, a few species of the submersum group cannot be excluded in the Neogene of Europe. Material: About 30 specimens and fragments. Inventorized under Nos BK-5465, BK-5469, BK-5523 LAMIACEAE Lycopus cf. europaeus L. (PI. II, Fig. 16) Description: Fruits with marginal rim of spongy tissue, obovate in outline, about 1 mm long and 0.B0 mm wide, marginal rim up to 0.1B mm wide aro- und apical part of the fruit; hilum basal, large; fruit body with fine longitudinal surface structure. Remarks: One specimen found without a marginal rim. The fruit remains of Lycopus L. haye been recorded from the Pliocene and Quaternary (see Van der BURGH 1983, BU?EK et al. 1985), the findings in the Miocene are rare (Van der BURGH 1987). Material: 3 specimens. Inventorized under No BK-5464 LYTHRACEAE Decodon gibbosus (E.M. REI0) E.M. REID (PI. II, Fig. 12): Description: Seeds are very thick-walled, shortly wedge-like form, small, about 1 mm high and 0.9 to 1.1 mm wide (deep); dehiscent valve in one ol the specimens is indistinct (in the other one it is missing), with surface structure hardly visible; seed surface smooth. Remark: The species is well known from the Pannonian and Pontian of southern Moravia (KNOBLOCH 1981a). There is a great need of taxonomic revision of the fossil seeds of Oecodon GMELIN because a few rather similar species with the uncertain delimitation were described. Material: 2 seeds. Invetorlzed under Nos BK-5503, BK-5538 ACTINIDIACEAE Actinidia faveolata C. et E.M. REID (PI. IV, Fig. 6): 55