Szabó János szerk.: Fragmenta Mineralogica Et Palaentologica 26. 2008. (Budapest, 2008)

Shape — Wide-based, moderately high turbiniform shell. Whorls markedly convex, but without obvious angulation, however, more convex upper and moderately convex lower ("outer face") belts distinguishable, narrow ramp without clear abaxial boundary exists. Suture rather strongly impressed; peripher} 7 rounded-angular. Selenizone (—15 % of distance between sutures) situated somewhat below midwhorl on moderately convex outer face. Base convex and phaneromphalous. Peristome not preserved on lectotype but cross-section of inner ("columellar") lip seems slighdy thickened on Bakony Mts specimens, visible at funnel-shaped umbilicus. Sculpture — Juvenile shell bearing canccllate ornament of equally strong spiral and collabral cords. Sub­sequent whorls and base predominandy ornamented with spiral cords and threads, alternating. Spiral cords lacking from selenizone, only threads visible there. On post-juvenile whorls, threads and cords provide collabral sculpture. Latter arise from upper suture and weaken into threads well before reaching selenizone. Density and initial strength of these elements similar to those of spiral cords. Growth-lines mar­kedly prosocline above selenizone, distincdy opisthocline between selenizone and peripher} 7 , and opisthocyrt on base. Remarks — HÖRNES (1853) and STOLICZKA (1861) regarded this species belonging to Trochotoma then SZABÓ (1980) accepted this generic attribution. However, verification of the presence and the place of the exhalant opening (trema), the most important basis for the generic arrangement remained missing. Subsequent studies (SZABÓ 1984) in order to find a systematic place for XJrkutitoma (see below) resulted in the recognition that in lack of trema, the best morphological character to distinguish trochotomids from the similarly shaped pleurotomarioideans is the orientation of the col­labral ornament between the selenizone and the peripher} 7 . This is (strongly) prosocline (and sometimes feebly proso­cyrt) in the trochotomids but orthocline or opisthocline and prosocyrt in the pleurotomarioideans. In this species, the growth-lines between the selenizone and the peripher} 7 are markedly opisthocline; therefore its attribution to the Trochotomidae has been erroneous. In lack of a true angulation on the whorls the most probable sys­tematic place is widiin Teptomaria. However, the subgradate outline of the shell reminds also the shape of Anodomaria. BLAKE (1872) identified a species as Trochotoma stria­tum that were regarded by TATE (1876) a synonym of his new Sinemurian species, Pleurotomaria ohesula. Though he expressed some differences from Trochotoma striatum, HÖR­NES's species remained also in his synonym list with question mark. Based on TATE's (1876) Pl. X: 6 figure, P. ohesula has gradate shell with wide (true) ramp and different measurements, thus it must belong at least to another genus and species. Distribution — Hallstatt, Hierlatz Alpe, Upper Sinemurian (Oxynotum Zone); Eplény and Fenyveskút (near Lókút) (Bakony Mts), Upper Pliensbachian. Figure 44 — Leptomaria Striata (HÖRNES, 1853), lectotype. — 2 a-b: STOLICZKA'S (1861) figures from Tafel V (copy); A-C: "apertural" (A), back (B) and apical (C) views, xl; D—E: magnified views to show the ornament, X2.

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