S. Mahunka szerk.: Folia Entomologica Hungarica 60. (Budapest, 1999)

Group C: typical species are P. mêlas, C. scheidleri, and Carabus convexus. This group provides favourable habitats for Carabus coriaceus, C. nemoralis and Harpalus atratus. Group D: the most typical species are A. parallelepipedus and A. bombarda. This group a wide range of species occurred with moderate frequency such as P. oblongopunctatus, Abax ovalis, Carabus hortensis, Platyderus rufus, C. scheidleri and P. melanarius. A. parallelepipedus was a single species that occurred in a relative high number in all sites of the four groups. It was with increasing importance from groups A to D, in axis one. A. bombarda is similar to A. parallelepipedus, but it was not in group A. Results of multivariate analises show that 15 sites of PBR maintain heterogenous carabid assemblages, with altering abundance and dominance of species. This evidently shows influence of various habitats within the given sites (cf. Baguette 1993; Kádár and Szél 1993). Furthermore, the results of the present study indicated that the knowledge of ground beetle species distribution in Hungary is also fragmentary, as Heijerman el ah (1989) have demonstrated by Dutch carabids in Netherlands. There are only data on the presence of typically and/or character species of the above groups, and there is a lack of information on distribution or quantitative registration and their analysis (excluding e.g. Báldi and Ádám 1991). In several papers it was shown that carabid beetles are good indicators of site and site conservation (Eyre and Rushton 1989). The typical and/or abundant carabids sampled by standardized sampling method in PBR are possible bioindicator species in all likelihood. Further, detailed investigations are needed to clear the role of carabid species in PBR. Acknowledgement — The authors are grateful to the late Dr. Imre Loksa (Eötvös Loránd University, Budapest), who collected a part of the material. REFERENCES Arndt, E. (1991): Beitrag zur Kenntnis der Licinini-Larven (Coleoptera, carabidae). —Ent. Nachr. 35: 117-121. Baguette, M. (1993): Habitat selection of carabid beetle in deciduous woodlands of southern Belgium. —Pedobiologia 37: 365-378. Báldi, A. and Ádám, L. (1991): Habitat selection of ground-dwelling beetles during dolomitic succession. — Annls hist.-nat. Mus. natn. hung. 83: 245-251. Ball, G. E. (1992): The tribe Licinini (Coleoptera: Carabidae): A review of the genus-groups and of the species of selected genera. —J. N. Y. ent. Soc. 100: 325-380. Borcard, D. (1982): Etude des communautés de Carabidae (Coleoptera) dans quelques associations forestières de la région neuchâteloise: Aspects statistiques. — Bull. Soc. ent. Suisse 55: 169-179. Brandmayr, P. (1974): Auto- und synökologische Untersuchungen über die Carabiden zweier Vegetationseinheiten des Slowenischen Küstenlandes: das Carici (humilis)-Seslerietum juncifoliae und das Seslerio (autumnalis)-Fagetum (Coleoptera, Carabidae). — Acta entomol. Jugoslavica 10: 15-40. Brandmayr, P. (1983): The main axes of the coenoclinal continuum from macroptery to brachyptery in carabid communities of the temperate zone. —• In: Brandmayr, P., Den Boer, P. J. and Weber, F. (eds.): Ecology of carabids: The synthesis of field study and laboratory experiment. Centr. Agric. Publ. Doc. (PUDOC), Wageningen, pp. 147-169. Brandmayr, P. and Zetto Brandmayr, T. (1986): Food and feeding behaviour of some Licinus species (Coleoptera Carabidae Licinini). — Monitore zool. ital. (N.S.) 20: 171-181.