S. Mahunka szerk.: Folia Entomologica Hungarica 57. (Budapest, 1996)

unpaired receptaculum seminis in my recent publication (cf. Sziráki 1996). The split sclerotization of the ductus seminalis (which is partly incorporated into the bursa copula­trix) well developed. A probable apomorphy in the FEIG is the presence of two different postbursal accessory glands with a common opening. This seems to be a synapomorphic feature of the families Chrysopidae, Hemerobiidae and Polystoechotidae (cf. Sziráki 1996). As regards the larval antenna of Polystoechotes punctatus (Fabricius, 1793), its structure (in the case of the hitherto known first instar) (Withycombe 1925) is the same, as in third instar of Dilaridae (Minter 1992) within the Osmyloidea. In Chrysopidae the FEIG (e.g., Principi 1977) shows a derived structure, because here the ductus seminalis is separated from the bursa copulatrix. Usually near to its proximal end this duct modified as "spermatheca" with strong sclerotization. At least a part of this sclerotization may be homologous with the split sclerotization in the plesiomorphic types of the ductus seminalis incorporated into the bursa. On the dorsal-, or antero-dorsal part of the bursa copulatrix there is a pair of bursal (accessorial) gland. This type of glands is also present in some osmylids (Adams 1969), and in psychopsids as well (see later); consequently, it must be regarded as a plesiomorphic character in Chrysopidae. The basic structure of the larval antennae of Chrysopidae is similar to that of Osmy­loidea. The four antennái section of the examined Mallada sp. (Figs 9-10) probable are homologous with those of Sisyra fuscata (Fabricius, 1793) (Figs 5-6). However, the lar­val antenna of Chrysopidae has an important apomorphy, namely: the flagellum is gradually and uniformly tapering towards its pointed apex. In Hemerobiidae (similarly as in Chrysopidae) also a part of the ductus seminalis is regarded conventionally as receptaculum seminis, but here there is a paired organ with a usually long, thin duct which is connected to the proximo-ventral part of bursa copula­trix. It is regarded to be homologous with the bursal glands of Chrysopidae (and thus with those of Osmylidae) (Monserrat 1990), however its shape and position well agree with the paired spermatheca of Ithonidae, Osmylidae, Polystoechotidae, etc. Conse­quently, I think it to be homologous with the plesiomorphic type of receptaculum semi­nis. (On the other hand, it is possible that this organ already have lost the spermatheca­function in the family Hemerobiidae.) The duct between bursa copulatrix and median oviduct usually is separated from the bursa, but recently was described a new hemero­biid species (Oswald 1994Ű) with duct shortly attached to the ventral side of bursa and with a strongly sclerotized section in its proximal part. The latter obviously is homolo­gous with the plesiomorphic split sclerotization in the fertilization canal of Ithonidae and some other families. As regards the larval antenna of hemerobiids, it virtually has the same structure, as that of chrysopids, including the gradually tapering shape towards the pointed apex. This feature may be considered as a synapomorphy of Chrysopidae and Hemerobiidae. The phylogenetic position of the Psychopsidae seems to be uncertain. These lace­wings are regarded by New (1991) as a family of Hemerobioidea, while by Henry (1978) as that of Myrmeleontoidea. Hitherto any sure synapomorphies were not found neither in FEIG, nor in larval antenna of this family and Hemerobiidae + Crysopidae + Polystoechotidae, or of Psychopsidae and the four myrmeleontoid families. In FEIG the paired spermatheca absent (cf. Sziráki 1996). This is an apomorphic character status, but it also developed independently in Chrysopidae and in Myrmeleontoidea. Paired bursal glands usually present (Oswald 19946) — as in chrysopids also —, but it is a plesiomor­phic feature. Ductus seminalis partly incorporated into the bursa copulatrix, and has a very strong split sclerotization.

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