S. Mahunka szerk.: Folia Entomologica Hungarica 57. (Budapest, 1996)

CONIOPTERYGOIDEA Coniopterygidae is the single family within this superfamily. It is a well separated, and presumably archaic taxon with many autapomorphies (Meinander 1972), however at present it seems to be uncertain, whether its separation occurred before or after that of Osmyloidea. As regards the FEIG, in case of subfamilies Brucheiserinae and Aleuropteryginae the separated, membraneous spermatheca is clearly-, while in subfamily Coniopteryginae it presumably is homologous with the plesiomorphic paired spermatheca of ithonids, osmylids, etc., but represents a derived character status. In the hitherto studied Brucheiserinae and Aleuropteryginae taxa the ductus seminalis partly incorporated into the bursa copulatrix and has a strong, split sclerotization as in Ithonidae, while in Coniopteryginae the fertilization canal runs from the receptaculum seminis to the common oviduct without touching the bursa copulatrix (FEIG type II and III, respectively) (cf. Sziráki 1996). In this family the larval antenna also shows a peculiar derived structure; it is only two segmented, altering from the plesiomorphic five- or four segmented larval antennae of Ithonidae, or of Megaloptera and Raphidioptera. OSMYLOIDEA FEIG of the four families counted to here (Osmylidae, Dilaridae, Sisyridae, Nevrorthidae) is similar to that of Ithonidae, but the ductus seminalis (which partly incorporated into the ventral side of the bursa copulatrix) is without the special split sclerotization mentioned above. This form of ductus seminalis may be regarded here as a synapomorphic character status within the Neuroptera. Some other suspected synapomorphies of the families of this group were detailed in a recent paper (cf. Sziráki 1996). In the osmyloid families the larval antennae show a rather modified structure (comparing those to the same organs of Ithoniidae). In basic case this type of antenna consists of four parts. I think to be reasonable the opinion of Wundt (1961), who investigated the musculature and innervation of the antenna of Osmylus fulvicephalus (Scopoli, 1763), and regarded the first segment (basal part) as a merged scapus and pedicel because of the presence of a chordotonal organ here. Consequently, the remaining parts of the antenna (in present paper: apical-, subapical- and middle part) must be regarded as flagellum. An other possibility: the basal part represents only the scapus, while pedicel merged with the first flagellar segment (Rousset 1966), i.e., it incorporated into the "middle part" of the antenna. The segmentation of this middle part often is incomplete (Osmylidae, Dilaridae, Chrysopidae, Hemerobiidae), and the number of the segments (or transverse grooves) within this part are increasing with the development of the larvae (Withycombe 1925, Minier 1992). Besides, in the families Osmylidae and Dilaridae at the apical end of the middle part there is a stout conic bristle, which may be a basiconic olfactory sensillum. All of the three parts of flagellum are distinct in Sisyridae (Figs 5-6). Here the thin, minute apical segment at the apex has a relatively long, straight bristle, while laterally a smaller one. The subapical part slightly swollen with a curved apical bristle. The middle part (i.e., the longest one) consists of about 10 segments in the third, and only 2-3 in the first larval instar. Its apical end is slightly swollen and also has a strong, curved bristle (Weissmair and Waringer 1994).