S. Mahunka szerk.: Folia Entomologica Hungarica 55. (Budapest, 1994)

Bubalonthophagus gen. n. Type species: Onthophagus furciceps Marséul, 1869 (present designation). Body surface finely microreticulated, weakly shining. Frontal ridge distinct in both sexes. Vertical ridge prominent, with a median tooth and with long, arcuate horn at both sides (male) or with short horn at both sides (female). Anterior part of pronotum with double median knob in both sexes, lateral knobs indistinct. Pronotal punctuation granulated. Citellonthophagus gen. n. Type species: Scarabaeus vitulus Fabricius, 1776 (present designation). Body surface finely microreticulated, weakly shining, almost dull. Frontal ridge distinct in both sexes. Vertical ridge prominent, with a short, erect horn at both sides (male) or without horns (female). Anterior part of pronotum with double median knob and with simple lateral knobs in both sexes. Pronotal punctuation granulated. Troglonthophagus gen. n. Type species: Scarabaeus semicornis Panzer, 1798 (present designation). Body surface finely microreticulated, weakly shining, almost dull. Frontal ridge distinct in both sexes. Vertical ridge prominent, upper margin deeply (male) or shallowly excised (female). Anterior part of pronotum with double median knob and with simple lateral knobs in both sexes. Pronotal punctuation granulated. Kisonthophagus gen. n. Type species: Scarabaeus ovatus Linnaeus, 1767 (present designation). Body surface finely microreticulated, weakly shining, almost dull. Frontal ridge indistinct in male, distinct in female. Vertical ridge simple in both sexes, more prominent in male. Anterior part of pronotum without knobs. Pronotal punctuation granulated. Lucidonthophagus gen. n. Type species: Copris lucidus Illiger, 1800 (present designation). Body surface polished, strongly shining. Frontal ridge distinct in both sexes. Vertical ridge prominent, with a short, erect horn at the middle (male) or with short horn at both sides (female). Anterior part of pronotum with double median knob and with simple lateral knobs in both sexes. Pronotal punctuation granulated at anterior and anteriolateral region, discal punctuation simple, very fine.