O. Gy. Dely szerk.: Vertebrata Hungarica 23. (Budapest, 1989)

Topál, Gy.: Tertiary and Early Quaternary remains of Corynorhinus and Plecotus from Hungary (Mammalia, Chiroptera) 33-55. o.

Plecotus (Plecotus) pliocaenicus Topái, 1988 This is one of the earliest Plecotus (Plecotus) species in Hungary. It was found in the Estramontian horizon of the Lower Pliocene of the Osztramos Locality 9. Being a greatsized Plecotus, in some respects related to Plecotus (Plecotus) austriacus (Fischer, 1829), and it has greater measurements than those of Plecotus (Plecotus) abeli Wettstein, 1923. Holotype: A right mandibular fragment (see Plate 11, Figs 9, 10) with I/1-I/2-I/3C-P/2-P/3-P/4-M/1 Reg. No. V. 86. 14, deposited in the collection of the Palaeontological Department, Hungarian Natural History Museum, Budapest. Ser. No. 111. Measurements of the Holotype: LCP 2. 56, LCPA2.41, LCL 0. 83, LCW0.92, LPTL 0. 59, LPTW0.77, LTPL 0.47, LTPW0.65, LPFL 0. 77, LPFW0.72, LMOL 1.50, LMOW 1.09, HMUM 1. 54. Other material and selected paratype : Reg. No. V. 86. 16., ser. No. 86, damaged rostrum with damaged right P4/, intact M1/-M2/ and alveoli of II/P2/, M3/ and alveoli of Ieftll/-P2/; Reg. No. of maxillary pieces V. 86. 17: ser. No. 87, right maxillary fragment with P4/- Ml/; ser. No. 88, right maxillary fragment with M2/-M3/; ser. No. 89, right maxillary fragment with Ml/; ser. No. 91, left maxillary fragment with C-P2/-P4/-Ml/ and alveoli of II/12/; ser. No. 92, left maxillary fragment with M1/-M2/ and alveole of M3/; ser. No. 102, left maxillary fragment with C, P4/-Ml/M2/ and alveoli of II/12/, P2/; ser. No. 10 3, left maxillary fragment with P4/-Ml/ and alveoli of II/-P2/; ser. No. 10 5, right maxillary fragment with P4/-M1/ and alveoli of II/P2/, M2/; ser. No. 106, right maxillary fragment with Ml/-M2/; ser. No. 107, right maxillary fragment with M2/; ser. No. 116, left maxillary fragment with P4/- Ml/; ser. No. 117, left Ml/; Reg. No. of mandibular pieces V. 86. 18. : ser. No. 93, left mandibular fragment with P/4-M/l; ser. No. 95, left mandibular fragment with alveoli of I/l-P/4; ser. No. 97, right mandibular fragment with P/2-P/3-P/4M/l-M/2 trigonid; ser. No. 98, right mandibular fragment with P/4 and alveoli of I/l-P/3; ser. No. 100, right mandibular fragment with P/4 and alveoli of C-P/3; ser. No. 110, right mandible with missing articular process, with P/4-M/l, damaged M/2-M/3 and alveoli of I/l-P/3; ser. No. 112, right mandibular fragment with P/4-M/1-M/2 and alveoli of I/2-P/3; ser. No. 65, left mandibular fragment with no teeth but alveoli of C-M/l; ser. No. 75, right mandibular fragment with alveoli of C-M/2; a selected paratype (see Plate II, Fig. 11), Reg. No. V.86. 15., ser. No. 90, left maxillary fragment with C-P2/-P4/-Ml/-M2/M3/. Measurements of the selected paratype: MHP 1. 86, MHM 1. 28, AB 0. 32, UCM 5. 91, UCMA 5.66, UPM 4. 50, UPMA 4. 36, UCP 2. 72, UCPA 2. 45, UMM 3. 64, UMMA 3. 18, UCL 1. 09, UCW 0. 95, UPTL 0. 50, U

PTW 0. 68, UPFL 1. 36, UPFW 0. 92, UMOL 1. 54, UMOW 1. 77, UMTL 1. 54, UMTW 1.9 5, UTML 0. 68, UTMW 1. 72. Measurements: including those of holotype and selected paratype, minimum and maximum values (number of cases: mean in parentheses). IOB 4. 18, CCRA 3.99, BIN 2.03, DIN 2. 00, WAPE 1. 54, DAPE 1. 22, WRL 5. 45, MHP 1. 59-2. 0 3 (8: 1. 874), MHM 1. 13-1. 50 (7: 1. 277), AB 0. 16-0. 32 (4:0. 270), UCM 5. 91, UCMA 5. 35-5. 66 (3:5. 500), UPM 5. 40, UPMA 4. 36-4. 36 (3: 4. 360), UCP 2. 68-2. 86 (3:2. 7 53), UCPA 2. 45-2. 59 (6:2. 343), UMM 3. 64, UMMA 3. 18-3. 32 (3: 3. 240), UCL 1. 04-1. 09 (3: 1. 0 73), UCLA 0. 81-0.90 (3:0. 856), UCW 0. 95- 1. 00 (3:0. 9 66), UCWA 0. 59-0. 68 (3:0. 650), UPTL 0. 50-0. 5 5 (2:0. 52 5), UPTW 0. 61-0. 68 (2:0. 654), UPFL 1. 22-1. 50 (7: 1. 342), UPFW 0. 92- 1. 04 (7:1. 021), UMOL 1. 311. 54 (12: 1. 427), UMOW 1. 59- 1. 81 (12: 1. 702), UMTL 1. 42-1. 54 (7: 1. 481), UMTW 1. 72-1.95 (7:1. 865), UTML 0. 63-0. 68 (2:0. 650), UTMW 1. 72-1. 97 (2:1. 845), LCMA 6. 11, LPM 4. 68, LCP 2. 56, LCPA 2. 08-2. 45 (8:2.276), LMM 3.93, LMM A 3.77, LCL 0. 83, LCW 0.92, LPTL 0.59-0.59 (2:0. 590), LPTW 0. 72-0. 77 (2: 0. 745), LTPL 0. 45-0. 47 (2: 0. 458), LTPW 0. 630. 65 (2:0. 640), LPFL 0. 72-0.90 (7:0. 818), LPFW 0. 72-0. 81 (7:0. 774), LMOL 1.45-1. 54 (5:1.514), LMOW 1. 04- 1. 13 (5: 1. 092), LMTL1.54, LMTW 1.15, LTML1.09, LTMTR 0.9 5, LTMTA 0. 72, HMUM 1. 40- 1. 59 (6: 1. 520), H MB M 1. 45, HCP 3. 18. Comparisons and remarks. P. pliocaenicus from Osztramos Loc. 9 furnished a rather fragmentary material with almost all teeth (except II/ and 12/) represented but not in a complete upper or lower row. It seems to combine the size and some features of P. austriacus and some characters of P. auritus. The interorbital breadth (a single specimen) is exceedingly great, greater than that of any specimens used in this study. On the other hand, the C-C alveolar width of the rostrum of the species is smaller than that of P. austriacus.