O. G. Dely szerk.: Vertebrata Hungarica 15. (Budapest, 1974)
Topál, Gy.: The First Record of Megaderma in Hungary (Pliocene Sediments of Osztramos, Locality 10) 95-104. o.
strength of the oral portion of the mandibular ramus and the strength of the premolar row. In this diagram I included, besides Rh. grivensis , all available specimens of the small species from Osztramos, and three from Podlesice. The regression equations are as follows: for lissiensis : Y = 0, 308664 + 0, 6428 x for the Carpathian euryale: Y = 0, 512213 + 0, 6329 x for the Dalmatian euryale: Y = 0, 839954 + 0, 4119 x There are regressions between the two variables at the P = 1 % and 2 % levels in each group. Anyway, there are no statistical differences between these equations, since lis siensis /Carpathian euryale P = 90 %, Carpathian euryale/ Dalmatian euryale P = 40 %, lissiensis /Dalmatian euryale 30 %> P> 40 %. Other diagrams (not shown here) reveal a C-M3/P2-P4 correlation very similar to that of P2 length/Mi-M3 length, because of differences in P 2 length. The C-M3 length measurements, however, actually are the same in Rh.lissiensis and in Rh.euryale. Regarding the length of P4 Rh . grivensis seems near Rh. lissiensis and at a distance from all Podlesice and Osztramos specimens of the smaller species. Contrarily, the M4-M3 length of Rh. grivensis aproaches that of the small species, while - regarding the combined P2-P4 length of Rh. grivensis given by MEIN - it partly fits into the measurements of the small form and partly agrees with those of Rh. lissiensis from Osztramos. The distal portion of humerus is slightly smaller, but it seems comparatively more robust than that of Rh. euryale. The trochlea is rather narrow, the epitrochlear portion short, but dorso-ventrally wide. The processus styloides is a little more ventrally pointed and witha smallerknob than in Rh . euryale . Besides, both trochlea and condylus are dorso-ventrally more elongated, that is, disto-proximally shortened in the fossil species. The epicondylus is smaller and shorter in the fossil form. After analysing the scatter diagrams, one may conclude that Rh. grivensis might have been the common ancestral form of the two small fossil Rhinolophi in the Pliocene of Europe. Evolution presumabely bifurcated through a small form into Rh. hipposideros and through Rh. lissiensis into Rh. euryale. These intermediate forms clearly existed together in the second half of Pliocene of Osztramos localities 1 and 9. Although the differences between Rh. grivensis and both species of Pliocene Rhinolophi in this size group are not great, still we may regard neither one as a subspecies of Rh.grivensis. Else the coexistence of two subspecies were to be presumed. Therefore I regard Rhinoloph us lissiensis as a valid species. Myotis sp. Material. Fragmentary proximal portion of a left humerus. Remarks. At present no exact identification is possible. Size about Myotis daubentoni . Myotis cf. aemulus HELLER Material. Right M , right M^, right lower C. 102
