B. Papp szerk.: Studia Botanica Hungarica 39. 2008 (Budapest, 2008)

Erzberger, P.; Schröder, W.: The genus Schistidium (Grimmiaceae, Musci) in Hungary

Scbistidium papillosum Culm. (Figs 25, 26) Plants small to large, in upper part often reddish, forming lax or dense tufts or mats. Central strand absent or rarely narrow and indistinct. Hair-point 0-1.25 mm, thin, flexuose, from shortly to longly and widely decurrent, slightly to strongly spinulose-denticulate in lower part, distantly denticulate to smooth in upper part. Costa ± densely papillose on abaxial side (papillae 6-8 urn wide and 5-11 urn tall). Margins ± papillose-denticulate in upper part of leaf. Lamina unistratose or with bistratose spots in upper part, ± densely papillose on abaxial side (papillae 6-7(-9) urn wide and 5-8 urn tall), on the adaxial side scattered papillae particularly near the margins. Lamina cells incrassate, sinuose, 8-10 urn wide in upper, 8-11 wide in central and lower parts of leaf. K+ red. Sporophytes common, from deeply immersed to emergent. Urn dark reddish, widest at about middle, length/width ratio 1.5-1.9-2.3(-2.8) (BLOM 1996), in Hungarian material 1.81, n = 1. Exothecial cells transversely rectangular and square. Stomata 10-17 per urn. Peristome teeth (310-)330-500 urn, reddish, gradually narrowed into a long, fine point, entire or with few to several median or submarginal perforations. When growing in sufficiently sun-lit places, S. papillosum usually (but not always) develops red-coloured spots in the lamina of upper leaves, which are diagnostic. In other respects, e.g. papillosity, hair-point length, this species is highly variable. Habitat: In Scandinavia, S. papillosum grows on both acidic and basic rock (from granite to limestone, but becoming more calcifuge towards the south) and occurs in dense forests as well as at exposed sites in open pastures (BLOM 1996, WEIBULL 2006). It prefers acid rocks in Catalonia (CASAS et al. 2001). In Hungary, this species was only recently discovered during fieldwork of the first author, and is presently known from a single location in the Bükk Mts, in the valley Lök-völgy near Felsötárkány, where it grows in open vegetation on aleurolitic slate near the road, associated with 5. apocarpum and S. pruinosum. Vertical distribution: 445 m a.s.l. Specimen examined: Bükk Mts: 7988/4 Felsötárkány, Lök-völgy, aleurolite slate rocks at road, N 48° 01 ' 51.1 ", E 20° 28' 46.5", 445 m, 06.04.2007, leg. P. Erzberger and T. Pócs, herb. Erzberger, (B) 12097.