B. Papp szerk.: Studia Botanica Hungarica 38. 2007 (Budapest, 2007)

Németh, Csaba: Data on the distribution of some rare alpine-boreal bryophytes in the Vértes Mts (Hungary)

into two major parts, the areas north of the line dominated by limestone, and those to the south by dolomite (Fig. 1). The north facing steep rocky walls of the dolomite areas have preserved a number of glacial relict vascular plants (discussed as the "dolomite-phenomenon", ZÓLYOMI 1942). These microhabitats are also important réfugia for alpine-boreal bryophyte species (BOROS 1968). The tops of the rocky outcrops are usually covered by rocky grass communities, in most cases classified as "closed dolomite rocky grassland" (Festuco pallenti-Brometum pannoniéi Zólyomi 1958) with associating Primula auricula L. subsp. hungarica (Borb.) Soó, Carduusglaucus Baumg., Coronilla vaginalis Lam., Polygala amara L., Biscutella laevigata L., Daphne cneorum L., Phyteuma orbiculare L. The surrounding vegetation contains mixed karstic Fago-Ornetum Zólyomi (1950) 1958 on the lithosol or rendzina of the steep mountain sides below the rock walls, Primulo veris-Tilietum platyphyllae (Isépy 1968) Borhidi 1996 on dolomite scree, rocky forest with oaks (BÖLÖNI et al. 2003), beech forest (Daphno laureolae-Fagetum (Isépy 1970) Borhidi in Borhidi and Kevey 1996), with Carexalba Scop., Calamagrostis varia (Schrad.) Host, Ranunculus nemorosus DC, Aquilegia vulgaris L., and Sorbus aria L., Sorbus danubialis (Jav.) Kárp., Sorbus ulmifolia Kárp., and Sorbus aria s. I. x Sorbus torminalis (L.) Cr. (endemic microspecies) in the canopy layers, Moehringia muscosa L., Draba lasiocarpa Rochel, and Viola collina Bess, in wet places and smaller rock terraces. The more common bryophytes of the surveyed dolomite rocks and rock forests with wider ecological amplitude include Ctenidium molluscum (Hedw.) Mitt., Distichium capillaceum (Hedw.) Bruch et Schimp., Encalypta streptocarpa Hedw., Fissidens dubius P. Beauv., Gymnostomum aeruginosum Sm., Homalothecium sericeum (Hedw.) Schimp., Neckera crispa Hedw., Plagiochila porelloides (Nées) Lindenb., Reboulia hemisphaerica (L.) Raddi, Scapania calcicola (Arnell et J. Perss.) Ingham, and Tortella tortuosa (Hedw.) Limpr. Having ecological requirements similar to the bryophytes above, the rare lichen species Solorina saccata (L.) Ach. in all localities is found together with the bryophyte species discussed. Methods The fieldwork was planned so that I could systematically survey all potential habitats, rocky mountain sides of northern exposition, which might contain alpine-boreal bryophyte species. The determination of geological coordinates was implemented by Garmin Etrex Legend GPS. The precision of the coordinates is between 5-10 metres. The geographical names follow those of the regional topographical map, scale 1 : 10,000, the map of forestry management plan, scale 1 : 20,000, and the tourist map, scale 1 : 40,000. Processing of the map was rendered with EOV system, using ITR interactive map editor system. The nomenclature of the bryophytes follows ERZBERGER and PAPP (2004), the nomenclature of the vascular plants follows SIMON (2000), and the nomenclature of the habitats is based on the work of BORHIDI and SÁNTA (1999). Entries of occurrences in the Enumeration below are based on data from the bryophyte collection of the Department