B. Papp szerk.: Studia Botanica Hungarica 30-31. 1999-2000 (Budapest, 2000)

Erdei, Boglárka; Lesiak, Maria: A study of dispersed cuticles, fossil seeds and cones from Sarmatian (Upper Miocene) deposits of Sopron-Piusz puszta (W Hungary)

ranged. Length and width of stomata are 28.8-37.5 urn and 20-28.8 urn, respectively. Stomatal pore is 12.5 um long. Type of stomata seems to be anomocytic. The outer stomatal ledge is heavily thickened. Description (BP 99.507.1): Thin cuticle with isodiametric cells. The hardly recognizable anticlinal cell walls are straight and thickened. The length and width of the randomly orientated stomata are 22.5-25 urn and 15-18.8 urn, respectively. Their smaller size may be attributed to its strongly compressed state. Length of stomatal pore ranges between 6.3-12.5 urn. Type of stomata cannot be accurately defined due to preservation. Discussion: According to KVACEK et al. (1982) stomata seem to be anomocytic, however, since the lateral subsidiary cells are sunken, they should be defined as cryptolaterocytic. In Hungary, Buxus pliocenica fossils (with cuticles) were described from the Pliocene alginite flora of Gerce, as the one and only evergreen element of the Mixed Mesophytic Forest (FISCHER and HABLY 1991). In Eurasia, Buxus leaf fossils are known from Tertiary and Quaternary deposits, first of all from Central and Eastern Europe (Stare Gliwice, Moravská Nová Ves, KVACEK et al. 1982; Willershausen, STRAUS 1969), Turkey, Georgia and from Kazakhstan. Based on fossils from Tertiary deposits, two main European lineage are distinguishable (KVACEK et al. 1982). The more primitive and extinct lineage is that of Buxus egeriana Kvacek, Bûzek et Holy. The second one, Buxus pliocenica Sap. et Mar., characterised by smaller leaves, comes from younger sediments, from the Middle and Upper Miocene. Considering the ecological requirements of Buxus, most species favour rather open areas with at least 800 mm annual mean precipitation, but they are well adapted to a seasonal climate. Exceptionally, they enter semiarid to arid regions (KVACEK etal. 1982). ?Lauraceae, ?Helobiae Wackersdorfia dubia Peters 1963 (Figs 21-22) 1963 Wackersdorfia dubia Peters; PETERS, p. 9, Taf. 2, Figs 10-14. Material: BP 99.508.1. Description: A thick cuticle with presumably isodiametric cells. Anticlinal cell walls, if observable, are straight and heavily thickened. Some stomata are recognizable, but their type cannot be identified. Length and width of the randomly